Oryza sativa L.

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Graminae

Common names

Rice.

Description

An annual grass with erect culms 0.6-2 m tall usually with four to five tillers. Inflorescence a loose terminal panicle of perfect flowers; each panicle branch bearing a number of spikelets, each with a single floret. Each flower is surrounded by a lemma and palea at the base of which are two small glumes. The lemmas may be awnless or variously awned. The rice grain enclosed by the lemma and palea (hull) varies in size, texture and colour. Each panicle holds 100-150 seeds.

Distribution

Throughout the tropical world as a crop.

Season of growth

Summer.

Rainfall requirements

Rain-grown rice usually requires an annual rainfall in excess of 1 500 mm. Most of the world's rice is grown under irrigation and water is supplied as required. The amount may be in the vicinity of 7-9 million litres per hectare.

Drought tolerance

It has little drought tolerance.

Soil requirements

Upland varieties need fairly good drainage but japonica varieties need a heavy, relatively impervious soil. The pH is generally not important, as rice has the capacity to neutralize the soil on which it is growing.

Sowing methods

The seed may be drilled into dry land or sown in nurseries and the seedlings later transplanted into a wet paddy-field. Also, seed may be pre-germinated and broadcast into the mud in the paddy-field.

Number of seeds per kg.

About 40 000.

Tolerance to herbicides

Rice tolerates the herbicides used to control its main grassy weed pest, Echinochloa crus-galli (barnyard or water grass). It will tolerate the use of 2,4-D amine at 1-1.5 kg AI/ha and 2,4,5-T amine at 1-1.5 kg AI/ha at the late tillering stage before panicle initiation for the control of broad-leaved weeds.

Dry-matter and green-matter yields

Yields of paddy straw after harvest varies considerably. A yield of 5 000-15 000 kg/ha may be expected.

Suitability for hay and silage

Practically all the paddy straw from rice crops in the tropics is conserved as hay for animal feeding and is usually stacked around poles in the house compound. Medling, (1972) under high rainfall conditions (3 997 mm) at Gualaca, Panama, found that baling the straw with large roller balers was effective and that the straw bales would stay in the field without major deterioration, while grass hay (Hyparrhenia rufa and other grasses) suffered serious deterioration.

Toxicity

Excess feeding of rice straw leads to toxicity because of high oxalates, which will bind the calcium in the diet. This effect can be reduced by soaking the straw in water or neutralizing it with a weak solution of calcium carbonate or calcium hydroxide (Göhl, 1975).

Seed yield

Japonica varieties, 4 000-4 500 kg/ha; indica, 1 000-1 500 kg/ha of milled rice.

Cultivars

There are numerous cultivars, each suited to a particular ecological niche.

Diseases

Numerous diseases occur, affecting the leaves for use as straw for feeding animals. The more important ones are blast, caused by Piricularia oryzae, leading to brown and shrivelled leaves; brown spot, caused by Cochliobolus miyabeanus; and narrow brown leaf spot, caused by Cercospora oryzae.

Optimum temperature for growth

A mean temperature above 21°C.

Frost tolerance

It will not stand frosts.

Maximum germination and quality required for sale

70 percent germinable seeds, 98.8 percent purity in Queensland.

Pests

Army-worms and grasshoppers cause major leaf damage, and stem borers affect the stalk.

Palatability

The hay is fairly palatable, but not nutritious enough for maintenance.

Response to photoperiod

It flowers over short and medium day lengths (Evans, Wardlaw & Williams, 1964).

Chemical analysis and digestibility

Göhl (1975) records analyses of vegetative material.

Natural habitat

Swampy areas.

Tolerance to flooding

The japonica or paddy varieties are aquatic and grow in water at various depths. Deep water varieties will grow in up to 6 m of water. Upland or indica varieties are not adapted to flooding.

Fertilizer requirements

In paddy rice, land preparation for planting usually involves some incorporation of organic matter, either from a previous grass/ legume pasture, green manure crop or from plants cut and transported to the field. A basic phosphorus and potash dressing may be required but nitrogen fertilization is a main determinant of yield. Half the nitrogen may be applied at transplanting with the remainder at ear initiation, and the application may be 150-250 kg/ha.

Genetics and reproduction

2n=24 (Fedorov, 1974). Some tetraploid (2n=48) exist, but the diploid are more numerous.

Seed production and harvesting

The rice is either hand harvested and later threshed or mechanically harvested with combines and hulled and cleaned.

Economics

Rice is one of the world's two major human food crops, the other being wheat. Rice straw is retained for feeding draught animals in most rice-producing countries in the tropics. Rice bran is fed to domestic animals when not required for human consumption in dry years. Paddy straw provides 80 percent of the organized roughage for India and a large part of the roughage for animals in other rice-producing countries where draught animals are used.

Animal production

The inclusion of urea and molasses improved the voluntary intake of paddy straw from 1.5 to 2.1 kg/100 kg live weight and dry-matter intake was 1.9-1.98 kg/100 kg live weight. Molasses-urea also increased digestibility coefficients of crude fibre and crude protein. This supplement increased body weight from 3.5 to 5.5 kg per head in 100 days. Paddy straw plus 1 percent urea and 18-20 percent molasses on a dry- matter basis is maintenance (Singh & Kushwaha, 1974). Sunn hemp (Crotalaria juncea) hay, with 12.7 percent digestible crude protein and 65 percent total digestible nutrients, provided maintenance for bullocks as a supplement to paddy straw in the proportions of 1:3 and 1:2. The bullocks consumed 1.952.11 kg DM/50 kg live weight (Ready & Murty, 1962).

Further reading

Chandraratna, 1963; Finlay, 1975.

Dormancy

The seeds of indica (upland) varieties have post-harvest dormancy for up to three months, which can be broken by storage at 8°C under very moist conditions immediately after harvest. The seeds of japonica varieties have no dormancy period.

Tolerance to salinity

Its extensive root system not only loosens the soil and renders it more permeable, to facilitate leaching, but also creates an acidic environment which reduces pH values in alkaline soils to neutrality. The improved cultivars 'IR-8' 'IR-8-68' and 'Jaya' exhibit high tolerance to salinity as do the tall indica cultivars 'Jhona 349', 'Damodor' and 'MCM' (Yadav, 1975).

Land preparation for planting

Upland rice and large-scale paddy cultivation where the seed is drilled prior to flooding need a well-prepared seed-bed, as for wheat. Paddy rice fields are usually ploughed and harrowed several times in the wet state as a "mudding-up" operation which incorporates organic matter and levels the land.