Bothriochloa insculpta (A. Rich.) A. Camus

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Graminae

Synonyms

Amphilophis insculpta (A. Rich.) Stapf.

Common names

Sweet pitted grass (East Africa), pinhole grass (Zimbabwe and South Africa), stippel grass (South Africa), creeping blue grass (Australia).

Description

A stoloniferous perennial, 30-100 cm high. Sessile spikelets shiny, shallowly grooved below the pit and glabrous, rarely with the margins finely hairy, pedicelled spikelets with three pits, rarely with one (Napper, 1965). It resembles Bothriochloa bladhii and B. ewartiana ­ all three have scented leaves and the scent persists in the hay. Creeping blue grass can be distinguished by its conspicuously hairy nodes, the reddish-purple colouring of the exposed stem portions and the development of creeping, as well as upright, stems. It differs from Dichanthium aristatum and D. annulatum in being scented, and having a pit on one side of the seed hull, and from D. aristatum in also having no hairs at the base of the branches (Bisset & Graham, 1978). Leaves, stems and seed-heads are aromatic and the aroma persists in stored hay.

Distribution

Kenya, Uganda, Zaire, southern and north-eastern tropical Africa. Common in Madras, the Nilgiris and Pulneys in India. Introduced to Australia as CPI 2695 in 1931 as Amphilopis (syn. Bothriochloa) glabra (Bisset, 1978).

Season of growth

Summer.

Altitude range

Sea-level to 2 000 m.

Rainfall requirements

The lower yearly rainfall limit is about 500 mm and its range generally 500-800 mm.

Drought tolerance

It is drought tolerant.

Soil requirements

It does very well on black clay soils but also on loams and clay loams, and will vegetate on eroded clays, scalded areas and puggy soils.

Ability to spread naturally

It spreads gradually ­ in one case 1 m on each side of a planted row in two years (Bisset & Graham, 1978).P

Land preparation for establishment

Establish on a well-prepared seed-bed, or in the ashes of a burn. Oversowing may give reasonable results.

Sowing methods

Surface sow and lightly roll or harrow; broadcast on to a scrub burn; broadcasting into barley stubble and brigalow is effective. In aerial sowing push the seed out of the hopper by hand (Bisset & Graham, 1978).

Sowing depth and cover

Not deeper than 1 cm.

Sowing time and rate

Sow from November to January (summer) at 2 kg/ha. For soil conservation sow at 9 kg/ha.

Number of seeds per kg.

About 1 210 000 fertile spikelets with one seed in each.

Seed treatment before planting

A treatment to remove awns from the seeds would assist sowing. A "de-awner" used for Stylosanthes humilis is successful with B. insculpta seed.

Seedling vigour

Good.

Vigour of growth and growth rhythm

A summer-growing perennial, it continues to grow into winter until cut by frost, but is slow in coming away in the spring in comparison with Rhodes grass and green panic.

Response to defoliation

Heavy grazing by hippopotamuses leads to dominance of Heteropogon contortus around Lake Edward, Uganda, and still heavier use will lead to poor Sporobolus and Aristida weeds (Heady, 1966).

Grazing management

Graze when the runners are well developed. Trampling by stock will develop the ground cover. Graze well during summer to prevent if from becoming stemmy. In pure stands or with Stylosanthes humilis keep it grazed to 10-15 cm; with siratro let the pasture reach 30 cm (Bisset & Graham, 1978).

Response to fire

Good. An annual burn is necessary if grazing is not heavy.

Dry-matter and green-matter yields

No records of yields have been found.

Suitability for hay and silage

It has been made into hay before the grass becomes stemmy. The hay has been eaten by horses and beef cattle but rejected by dairy cattle (used to better material) and sheep. With siratro its palatability is better. The hay is aromatic.

Value as a standover or deferred feed

It is useful as low quality roughage but is rather stemmy.

Toxicity

The scent of the flower heads will not cause taint in milk if eaten, though it deters cattle when more palatable feed is available.

Cultivars

The only registered cultivar is cv. Hatch ­ after Garney Hatch of The Caves, Rockhampton, Queensland, who sponsored its early acceptance. It is described above.

Diseases

In a wet autumn a leaf rust (Puccinia duthiae) can reduce seed yields.

Main attributes

Its ability to compete with native grasses on forest country without nitrogen fertilizer and to invade Heteropogon contortus pastures. Its hardiness and ease of establishment.

Main deficiencies

It lacks a little in palatability because of aromatic leaves.

Frost tolerance

Good. It has some winter hardiness.

Response to light

It does not grow well in shade.

Ability to compete with weeds

Good.

Maximum germination and quality required for sale

It is difficult to get more than 50 percent purity, though germinations up to 70 percent can be obtained. Seed germination declines rapidly after two years.

Pests

There are no major pests.

Palatability

It is reasonably palatable, much better than Heteropogon contortus, and is improved in association with siratro.

Response to photoperiod

It flowers in late April and in November in central Queensland (Bisset & Graham, 1978).

Natural habitat

Open bush and grassland, mainly in heavy textured black soils.

Tolerance to flooding

It will not thrive under waterlogged conditions.

Fertilizer requirements

Although able to persist in unfertilized ground it does respond well to nitrogen fertilizer.

Compatibility with other grasses and legumes

It will grow with native grasses and gradually dominate them. Where tried, it combined well with siratro and Stylosanthes humilis (Bisset & Graham, 1978).

Genetics and reproduction

2n=50, 60 (Fedorov, 1974).

Seed production and harvesting

It flowers twice a year in Rockhampton, Queensland, on the Tropic of Capricorn, in autumn and late spring. For seed the pasture is grazed till late summer and shut till seed ripens in late autumn. The crop is direct headed. Clean seed plots have been established in Queensland to prevent contamination with seed of Dichanthium aristatum (Angleton grass). Seed yield: 10- 30 kg/ha with up to 80 kg/ha per crop with nitrogen fertilizer and irrigation.

Economics

One of the few grasses suited to sowing in black self-mulching clays. It has been used with fair success in reseeding the medium rainfall areas of Baringo, Kenya.

Animal production

Around Rockhampton (rainfall 1 495 mm) on the Tropic of Capricorn, carrying capacity is one animal to 1.2 hectares.

Further reading

Bisset & Graham, 1978.

Dormancy

There is little dormancy.

Value for erosion control

It is useful for erosion control on self-mulching black clay slopes. It is one of the few suitable grasses as it establishes quickly and forms a good ground cover. The runners do not take root readily, but this is improved by prolonged wet weather, trampling or by soil cover, and using high seeding rates. The pastures must be grazed or burnt periodically.