Zea mays L.

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Graminae

Common names

Maize (Australia, United Kingdom), corn, sweet corn (United States), bok'olo (Ethiopia).

Description

A coarse annual, culms 60-80 cm high, straight, internodes cylindrical in the upper part, alternately grooved on the lower part with a bud in the groove. The stem is filled with pith. Leaf-blades broad. Has separate staminate (male) and pistillate (female) inflorescences. The staminate inflorescence is a tassel borne at the apex, the pistillate flowers occur as spikes (cobs) rising from axils of the lower leaves. The ovary develops a long style or silk which extends from the cob and receives the pollen from the tassel.

Distribution

Originated in Mexico or Central America, now pan- tropical; also grown as a summer crop in temperate Europe.

Season of growth

Summer.

Altitude range

The period of flowering and to maturity varies greatly in East Africa. Allan (1973) divides Kenya into four zones:
  • Zone A 
below 200 m. Lowland tropics with high maximum and minimum temperatures. Quick-maturing varieties flower in two months and mature in four.
  • Zone B 
200-1 200 m. Most of these areas have low rainfalls and little maize is grown. In Australia most of the maize is grown from sea-level to 500 m.
  • Zone C 
1 200-2 100 m. Contains over 90 percent of the maize grown in Kenya. The highest yields are regularly obtained in this area. The Kitale hybrids (prefixed by the number 6) flower in about three months and mature in six at 1 500 m. At 1 800 m, flowering is at 3.5 months and maturity at 7, while at 2 100 m, the figures are 4 and 8, respectively.
  • Zone D 
2 100-3 200 m. Little maize is grown above 2 000 m, as only long-term varieties can survive in such high altitudes. At Ol Joro Orok (2 400 m) the maize takes 6.5 months to flower and more than a year to mature.

Rainfall requirements

An annual rainfall of more than 500 mm is needed, with best yields usually in the 1 200-1 500 mm area; it is often an irrigated crop. Kitale experiments show that the more rainfall after five weeks' growth, the higher the yield.

Drought tolerance

It is fairly drought tolerant up to five weeks, but thereafter is very susceptible. Dry weather at pollination time seriously affects pollination and hence yields.

Soil requirements

It requires a well-drained, fertile soil. Alluvial loams, deep latosols and clay loams are preferred.

Land preparation for establishment

A deep (20 cm) friable seed-bed should be prepared, as maize is comparatively shallow rooted and needs loose soil in which the roots can forage.

Sowing methods

It is usually drilled in rows for grain and fodder, though it can be broadcast thickly for turning in as a green manure.

Sowing depth and cover

Sow at 7.5-10 cm depth and cover using a fined instrument, then compact with a following press wheel.

Sowing time and rate

Late spring to midsummer, at row spacings of 105135 cm, using 9-11 kg seed per hectare with populations from 25 000 to 70 000 plants per hectare.

Tolerance to herbicides

Pre-emergence treatment with atrazine or simazine at 1.5-2.5 kg/ha controls most weeds, while 2,4-D amine at 0.45-0.55 kg/ha used on crops 25 cm high is a useful post-emergence weed-killer.

Dry-matter and green-matter yields

Yields of 10-50 tonnes of green matter per hectare are obtained for silage.

Suitability for hay and silage

Maize is conserved as stover in the United States and southern Africa, but the main use of the whole plant is as silage. It makes probably the best silage of the grass family, with heavy yields and high acceptability and without the need for additives. It is cut when the grain is full and glazed, in the medium dough stage.

Value as a standover or deferred feed

After the grain is harvested the residue usually contains 3.5-4.0 percent crude protein; in a drought year it can be as high as 9 percent. The protein level usually does not decline as it stands in the field. Moisture content at harvest is 40-50 percent in the United States, too low to make silage. The digestibility is 40-50 percent at harvest, but falls to 3638 percent after 40-60 days in the field. In the United States, good practice is to graze 25-35 percent immediately after harvest with one beast to 0.8-0.9 ha, and then harvest the rest for winter storage (Corah, 1979). In Kenya, it is usually dried in the field for several weeks after maturity (Acland, 1971).

Seed yield

1-4 tonnes of grain per hectare, which should be stored at 14 percent moisture or less. Minimum germination and quality required for commercial sale. 80 percent germinable seed and 98.6 percent purity in Queensland.

Cultivars

Numerous cultivars are available throughout the world and contact should be made with local extension officers to ascertain what is the current preference to specific conditions. Most countries have bred their own cultivars to suit their varying conditions.
Dent maize (Zea mays indentata) is the main variety grown commercially for grain and fodder, but there are other types such as pod corn (Zea mays tunicata), a curiosity; flour corn (Zea mays amylacea) for human consumption; flint corn (Zea mays indurata), preferred for the European market, with horny endosperm; sweet corn (Zea mays saccharata), used as a vegetable; pop corn (Zea mays everta), used as a snack food. High-lysine corn has been improved for human nutrition. Open pollinated varieties have been used for a long time but now most of the commercial dent maize is either a single cross or a double cross hybrid bred for special areas, soils and climatic conditions.

Diseases

Maize is subject to many diseases, chief of which are maize smut ( Ustilago maydis), head smut (Sphacelotheca reiliana), and various stalk and ear rots such as Gibberella and Diplodia. They have been overcome by chemical seed treatment or by breeding resistant varieties.

Optimum temperature for growth

Peak germination was at 20-30°C and growth at 18-21°C (Hughes, 1979).p

Minimum temperature for growth

8.7°C for Kitale hybrids in Kenya.

Frost tolerance

It is very susceptible to frosts.

Latitudinal limits

It has a wide range, from 58°N in Canada to 40°S.i

Ability to compete with weeds

It has poor competitive ability until the crop canopy has closed.

Pests

A variety of pests are encountered. Chinch bug (Blissus leucopterus) is a major pest in the United States. The corn ear worm (Heliothis armigera) is a problem in Australia.

Palatability

Excellent for all green matter.

Response to photoperiod

Some varieties are indifferent to day length, others require short days for flowering (Evans, Wardlaw & Williams, 1964).

Tolerance to flooding

Maize has no tolerance to flooding.

Fertilizer requirements

The needs for maize are best determined by soil tests. It generally requires a complete fertilizer, with heavy demands from about 40 days until maturity. Zinc deficiency causes leaf chlorosis and can easily be overcome by the use of zinc sulphate.

Compatibility with other grasses and legumes

To improve the nutritive value of forage, legumes such as cowpea can be planted between rows of maize. Lablab purpureus is used in Brazil. Grass seed can be undersown in maize as a means of establishment.

Genetics and reproduction

2n=20, 21, 22, 24 (Fedorov, 1974).

Seed production and harvesting

Maize matures in 80-120 days according to variety, and is either hand-picked and later threshed or is harvested with a combine.

Economics

Maize is a major human food grain in Africa and in eastern Indonesia. Some 85 percent of the maize crop in the United States is fed to livestock as grain and silage.

Further reading

Martin & Leonard, 1959.

Dormancy

There is little dormancy in maize seed.

Value for erosion control

As maize is usually a row crop and has a poorly developed root system, crops are very susceptible to erosion.

Tolerance to salinity

Maize gave maximum yields at ECe of 2 mmhos/cm, 50 percent at ECe 9 mmhos/cm and nil at 15.3 mmhos/cm (Maas & Hoffman, 1977). Further studies showed that in water cultures, or on mineral soils with surface irrigation and continuous leaching, the maximum salt concentration in the soil saturation extract that does not reduce maize yields is about 1 100 mg/l total dissolved salts (ECe - 1.7 dS/m). The maximum permissible salt concentration of irrigation water to sustain maize production is about 300 mg/l, an ECW of 0.45 dS/m (Hoffman et al., 1979).