Abstract
Oat
is grown in all parts of Tibet
and most of them are wild types of geno types. Extensive investigation
on oat genetic resources revealed that there are 11 different sub-species
of oat in Tibet.
This diverse biodiversity of oat crop and natural condition in Tibet
provides ample opportunity to promote oat as fodder and food crop. Research
is underway to select suitable varieties for fodder and food production
under different conditions. Crossbreeding between oat and wheat was undertaken
to develop a new variety which yielding, wide adaptation and resistance
to diseases.
Oat
Genetic Resources in Tibet
Oat
is an ancient crop, and it has been generally accepted that oat was originated
from China. Short growing
period and widely adapted to different types of soil, climates and ecological
condition are the main characteristics of oat. Though it is harsh condition
of the climate with the high altitude of Tibet, oat is growing
all parts of Tibet. Wild types of
oat are the commonly seen in Tibet.
In
1981, during the collection of oat genetic resources in Shannan Prefecture,
cultivated type of variety was not found but wild types of oat was found
widely in this area. It is distributed in the area where the altitude
is below 4500 masl. During this collection and inventory, 11 sub-species
of wild oat were reported. Many of these sub-species were transitional types of wild
oat from covered seed to naked seed. In 1982, not only many wild types
of oat were found but also many cultivated types of oat were found in
Changdu prefecture in eastern part of Tibet. It was reported that
these types of cultivated oats had been growing in the Jiangda county
of Tibet for many years. The main characteristics of these cultivated
types of oat were naked seed, but most of the small spike was close to
wild type with twisted long awn. With further evaluation and appraisal,
it was concluded that these new sub-species (mutations) were evolved directly
from primitive wild types of oat. In addition, in the BuoMi, ZuoGong County,
another types of wild oat, which was, tight spike with small spike handle
was found.
Main
achievements and status of research on oat genetic resources in Tibet
Extensive
investigation and collection
During 1981~1984, large-scale investigation and collection of oat genetic
sub-species of oat and its relative were found. Among them, 161 oaten
species were belonging to Avena fatua, 5 of them were H.virescens
of Helietotrichon von, and 4 of them were belonging to close relative
of oat.
Main Progresses and achievement
Along with the extensive collection and inventory of oat genetic resources,
extensive research on oat genetic resources had been carried out.
It was concluded that there were 11 different sub-species of oat in
Tibet and research on
the identification of cultivated types of oat were conducted.
Meanwhile,
research on crossbreeding of oat was carried out. Crossbreeding between
wild oat and winter wheat was done and new variety called FY-8 was developed,
which has characteristics of strong diseases resistance, widely adapted,
good quality of seed and well performed yield. At presently, many farmers
in Tibet are cultivating it.
Research on oat and durum wheat was also conducted to improve the wheat
seed quality.
Distribution of oat genetic resources in Tibet
Wild
oats in Tibet is locally called “ You Bu". It is widely distributed in Tibet. It is annual
crop with high stem and seed falls naturally when it is matured. Great
adaptability and strong reproductive make it very widely distributed wild
weed for farming in Tibet. It can be seen near by the field, in the farm field, along the road
and in some grassland field. Its wide spread is mainly human-induced because
people carry the seed with their grain and other farm products from one
place to another. However, naturally developed population of wild oat
has not been found in Tibet. With increasing of the altitude, the height of the oat straw reduces,
the length of the awn is shortened, seed covered increases, number of
the seed reduces, seed get smaller and spike become shorter. Wild oat,
which with hairiness at the outside shell of the seed, edge of the leaf
and plant stem are extremely widespread.
Most
of the Tibet oat is wild types of genotype. They are distributed
at the elevation between 2400-4500 meters. Cultivated
types of oat are often found in the counties of Jiangda, Gongjue in the
eastern parts of Tibet. The upper limit altitude is about 3000 meter.
Classification
and distribution of oat in Tibet
In
1753, oat was given the Latin name, as Avena as family under which,
there was four species were classified, such as A.nuda L., A.fatua
L., and A.sativa.L. and A.steilis L.
In 1958, three different species of the oat in China were identified. However, classification of oat species is still debating
and some of the Chinese scientist suggested that oat could be classified
into 18 different sub-species. Table 2 shows all the sub-species found
in Tibet. There were altogether
22 different types.
Characteristics of oat genetic
resources in Tibet
Common
wild oats
More
than 160 samples of common wild oats were examined and classified into
21 different types. Of which there are 5 types of them were light brown
seed, which has characteristics of loose inflorescence and hairiness of
outer shell of the first and the second floret. This
types accounts for 44%. There were 8 types which the seed color is gray,
which has characteristics of loose inflorescence and hairiness of outer
shell of the first and the second floret. This types accounts for 40%.
There was 1 types of them were brown seed, with loose inflorescence and
hairiness of outer shell of the first and the second floret. There were
8 samples were dark purple seed and 5 of them were transitional types
from covered seed to naked seed.
Cultivated type
Most
of the cultivated oats in Tibet are naked seed type. It was found in 1985 that the cultivated oat
in Tibet has the characteristics of each awn of the oat is twisted, the
bottom of the seed is smooth, stem high is 120-145 cm, the length is the
awn is about 20mm, the caryopsis is around 15 mm and it is loose inflorescence.
Experimental
test on the introduced varieties and local cultivars
During
1998-1999, 5 introduced oat varieties were test in Lhasa. Oat varieties
No. 1 and 2 were introduced from Inner Mongolia and No. 3, 4, and 5 were
introduced from Hebei Province. Two local cultivars Gongjue Oat and Jiangda
Oat were collected during early 1980s from the eastern part of Tibet in Gongjue County
and Jiangda County of Changdu Prefecture. All these varieties are naked
varieties. Initially these varieties were introduced and tested for food
purpose but later it was found that the straw of those oats were better
than barley and wheat for feed for animal. It was tried to introduce to
the local farmer to grow for fodder purpose but local farmer rejected
because in the central part of Tibet, traditionally
farmer think that oat is “wild-weed”. However, this experimental test
was an attempt to select better varieties for farmers for fodder and food.
It was sown on March 30th, germinated
on April 20th and tiller was occurred on May 20. The growing period of
these varieties were 132-150 days. Stem high ranges 120-145 cm, the length
of spike ranges between 20.7-25.4 cm. The 1000 kernel weight is ranging
from 21.9-20.9 g. The yields of these varieties range between 101.7-236.57kg/mu.
Meanwhile two local cultivars were also tested. They were sown on March
30th, germinated on April 16th and tillerring was occurred on May 10.
Comparing with those introduced varieties, the germinating date and tillerring
date of local varieties are 4 days and 10 days earlier, respectively.
The growing period of these varieties were 110-127 days. Stem high ranges
107-132 cm. Local varieties are earlier matured and higher straw than
those introduced oat varieties. However, the yields are much less than
the introduced varieties. It ranges from 89.1 – 127 kgs/ mu. Table 1
shows the main result of the test.
Table
1: Main Agronomic Characteristics of Introduced Oat Varieties and Local
Cultivars
| Oat
cultivars |
Growing
period (day) |
Stem
high
(Cm) |
Length
of spike (Cm) |
1000
kernel weight (g) |
Yields
(kg/ha) |
| No.1 |
145 |
174.9 |
|
21.7 |
3548.55 |
| No.2 |
132 |
68.2 |
20.7 |
20.9 |
1525.5 |
| No.3 |
139 |
100.8 |
22.1 |
21.4 |
3282 |
| No.4 |
137 |
132.1 |
21.9 |
21.6 |
2845.5 |
| No.5 |
150 |
194.9 |
24.5 |
21.9 |
3312 |
| GongJue
oat |
127 |
132.1 |
23.2 |
15.8 |
1905 |
| JiangDa
oat |
118 |
118.9 |
21.2 |
14.7 |
1336.5. |
Experimental
Test on comparing spring wheat, the introduced oat varieties and durum
wheat in Tibet
After
first year test in Lhasa, three oat varieties such as No. 1, 2, and 3
perform good yield in Lhasa. In order to further select the better cultivars
and to use its potential in Tibet, further test
on these varieties were carried out. Another aim of this experiment is
to see the quality of oat seed through comparing with common wheat and
durum wheat.
Durum
wheat is rarely grown in Tibet. However, the quality of grain of dominated common wheat varieties
is very unfavorable to process into bread and noodle. The main problem
lies here is that the protein content of the winter wheat is very low.
In most of the area, traditional spring wheat have been replaced by high-yielding
winter wheat cultivars. Durum wheat was also introduced to Tibet. New oat varieties
were also introduced in recent year. Testing on comparing wheat, introduced
oat and durum wheat were then conducted. It was also aimed at improving
the wheat grain quality through cross-breeding and mixing the grain into
wheat grain so that the overall protein content will be increased. This
attempt may not be ideal but protein content in oat seed and durum wheat
seed are tending to be much higher than that of winter wheat. Meanwhile,
direct use of oat for food and fodder and durum wheat for noodle in Tibet was also set as
one of the objective.
Yield
performances
Table
3 shows the main result of the test of 3 oat varieties, 5-durum wheat
and 2 high-yielding spring wheat cultivars. Yield of all cultivars range
from 101.7~236.7 kg/mu. Among all the varieties of three different crop,
oat variety No. 2 Yielded about 236.7 kgs/ mus, which was the highest
among all varieties tested and it was 78.5% higher that the yield of spring
wheat variety – Zangchun No. 6 and 18.9% higher that the yield of Rinong
No. 23. The yield of the oat variety No 1 was about 234.33 kg/mu that
was 76.6% and 17.8% higher than Zangchun No. 6 and Rinong No 23, respectively.
Among the three durum wheat varieties such as No. 291, No. 281, No. 294
and No. 266 Produced 7 – 39% higher yield that the wheat variety Zangchun
No 6 and 23- 49 % higher yield than Rinong No 23.
Analysis
of variances on yields of these varieties was done. It indicated that
there is no apparent variance among the replicates but there are large
variances among different varieties. T-test at 0.05 level and 0.01 levels
showed that the differentiation of the yield among different crops and
different varieties are considerably large. Particularly the yield of
oat variety No. 2 and No.1 are significantly higher that most of the varieties
of common wheat and durum wheat at 0.05 level. Table 4 and 5 shows result
of the analysis of variances of yield and t-test of significance of variation
among the yield.
Agronomic
characteristics
Growing period All
the varieties were sown on the April 20th, most of the varieties matured
within 135-150 days. The shortest growing period was the Oat No.2, which
was 132, and the common wheat variety-Zangchun No. 6 took longest to mature.
It was about 150 days.
Straw
height
The
height if the straw of all varieties tested ranged from 68-194.9 cm. The
oat No. 3 was the highest and the durum wheat No 5266 was the shortest.
Other varieties ranged from 72-190.2 cm.
Length
of the spike
The
length of the spike of oat varieties ranged from 20.7 – 25.4 cm. The length
of the spike of durum wheat ranged from 5.77 – 7.05 cm.
Number
of kernel in each spike
The
number of kernel in each spike of oat varieties ranges from 42 - 97.8.
The number of kernel in each spike of durum wheat ranges from 37.4 - 65.4.
1000
kernel weight
The
1000 kernel weight of oat is ranging from 21.9 - 20.9 g. The 1000 kernel
weight of durum wheat ranges from 28.6 - 44.4g
Potential
of oat as fodder and food crop as compared with common wheat and durum
wheat
The experimental tests were conducted during 1999 and 2000.
Compared the result of test in 1999,
in 2000 all the varieties were sown later to test the suitability of sowing
date of those varieties. The above experiment has shown that the
general agronomic characteristic of oat is better than durum wheat .The
average Yield of all cultivars of oat is higher than durum wheat. According
to this experimentation we can conclude that the adaptability of oat is
better than durum wheat. Except the No. 3 oat variety had serious lodging
at later stage that cause reduction in the yield of it. Oat variety No.
1 and No.2 performed very well in terms of yield and agronomic characteristics
and they are suitable for Tibetan condition for fodder production and
food production. Durum wheat needs good soil moisture content. When the
temperature is high in the spring and it makes lost in soil moisture content,
it affect the yield of durum wheat and common wheat. The yield of both
durum wheat and common wheat reduced when there was sever drought in springtime.
This is mainly because that low soil moisture content reduces the rate
of germination of wheat and eventually reduces the yield. But this does
normally not affect oat. The yields of oat in different year retain as
normal no matter how dry it was in the springtime to some extent. That
means these varieties of oat have higher resistance to drought than wheat.
Tibet is often very
dry in the springtime and if there is no irrigation, there will be major
failure in crop yield due to spring drought. In this case, wherever no
irrigation, oat can be potential crop to grow as food or fodder purpose.
However, there is little understanding on the crop oat. It has been locally
known as wild weed and little attempt had been made to promote this crop
for fodder and food. Two year experiments on testing different kinds of
cultivated oat varieties introduced from outside of Tibet has shown that
there is a great potential for oat to be highly adapted fodder crop and
food crop if it is properly processed into food item for the market.
Utilization
of oat genetic resources in Tibet
Unique
biodiversity of oat crop and natural condition in Tibet, provide great
opportunity to promote oat as fodder and food crop in Tibet.
Oat
as a fodder crop
Farmer
often harvests wild oat grown in the field of barley and wheat at different
stage. Farmer in most of the areas can easily recognize the young oat
plant among the barley and wheat crop. Fresh plant of oat is directly
fed to the animal. It is also dried and stored for feed sources during
the wintertime. In most of the cases, oat plant is harvested before maturing
so that the oat seed will not fall into the field. It is not only for
weeding purpose but also for collecting feed for livestock. Dried oat
straw is considered as high quality feed for livestock. In many case,
farmers use the collected wild-oat plant from the barley or wheat field
for animals like horse and milking cows. However, oat has not been widely
planted in large scale for fodder for animal. The main reason is that
farmer cannot collect enough seed from the wild oat for cultivation for
next year. In addition, they think that wild oat is weed and they do not
think it can be cultivated for feed. They know that it is good feed sources
and keep collecting from the other crop’s field.
In
recent year, scientists and agronomists tried to test new cultivated oat
varieties introduced from other provinces of China. Some of them
have selected suitable varieties for local condition. Selected varieties
were planted in different altitude ranges. For example, oat was grown
in Naqu prefecture and Damshong county of Lhasa. At the elevation of lower
than 4700 m, oat can grow very well for fodder. Because of its highly
drought resistances and adaptability to the harsh condition, oat crop
shows its great potential as fodder crop for higher altitude under the
pastoral and agro-pastoral system. One of the main constraints in developing
livestock in pastoral areas of Tibet is shortage in
feed during the wintertime. Those counties that practiced to grow oat
as fodder crop in high elevation have experienced that oat is very promising
fodder crop for feed production. In the areas where large area of cultivated
land is available but lacking of irrigation, oat as an alternative crop
for feed production has been recognized by many farmers and agricultural
developer. In Naidong County of Shannan Prefecture, large area of fodder
oat introduced from Qinghai provice was grown during 1996-1998. It was
aimed to produce feed for livestock production in upper region where sever
feed shortage is occurring during the wintertime. In that county, oat
was planted during the summer time and harvested in autumn and then processed
into silage or chopped into fine pieces that will be then for feed for
animals in upper region during winter season.
In
recent year, the government has focused on developing livestock in the
crop-dominated area in the central Tibet. However, lack of suitable feed crops and poor feed processing confine
further development livestock in these region. Barley straw is the main
source for feed for animal in many counties during the wintertime. But
the quality of the barley straw for animal feed is very poor. Livestock
production in many counties has been confined with shortage of feed during
the dry season. However, Tibet has vast land. There is scope for feed production. In order to boost
up livestock production in Tibet, both government
and researcher are looking for possibilities for promoting new feed crop.
Introduction and testing new fodder crop such as oat and corn is underway.
Fodder oat will be promoted in some region. But appropriate varsities
that adapt to the local condition are precondition for promoting oat crop
as fodder.
Another
concern of the local government is to develop livestock in pastoral area.
Overgrazing due to low carrying capacity of the rangeland in Tibet and increase in
the population of livestock is occurring. One of the main approaches taken
by the local government has been to promote adaptive and high-yielding
fodder crop for the region. Oat, in this sense, has its great advantage
of wide adaptation and cold resistance over the high and cold climate
condition in high altitude areas of Tibet.
Transformation
of agricultural development from subsistence farming system to market
oriented system is underway. Major policy to restructure the agricultural
production is adopting. Purely grain production based agricultural development
pattern is changing into a combination of cash crop, grain crop and fodder
crop. Oat as of the potential crop for high quality fodder will be promoted
in wider range of area wherever it is adapted to cultivate. Selection
of good varieties is essential. Above experiments suggested good variety
can be selected for most of the areas of Tibet through introduction of new varieties from outside of Tibet.
Oat
as food crop
Traditionally
oat was a food crop in eastern Tibet. But now it has
been no more there. High yield and good adaptability to the local condition,
oat has a potential to be food crop in Tibet as well. Beta-glucan
content in oat grain provides an opportunity for promoting oat as food
crop, particularly as a healthy food crop. Local people may not consume
oat as food, but it can be processed into healthy food for inner-land
of China. Our experiment
during the 1999-2000 showed that oat is well-adapted and high yielding
food crop if we select the right varieties and proper management.
Table
2: Agronomic Character of Oat in different altitude
| Altitude (m) |
< 3,000 |
3,000~3,500 |
3,500~4,000 |
>
4,000 |
| Straw
height (cm) |
120 |
105.8±18.0 |
102.5±17.8 |
100.3±18.5 |
| Length
of awn (mm) |
30±1 |
36±6 |
36±5 |
37±5 |
| Cover
of caryopsis (mm) |
25±2 |
23±4 |
23±3 |
21±2 |
| Width
of Cover of caryopsis (mm) |
7 |
57±0.6 |
59±0.8 |
61±0.8 |
| The
first cover of caryopsis
(mm) |
8.5±0.7 |
9.3±0.5 |
9.5±0.7 |
9.6±0.6 |
| The second cover of caryopsis (mm) |
8.5±0.7 |
9.7±0.9 |
10.3±0.8 |
10.6±1 |
Table
3.: Classification of oat in Tibet
|
Grain
colour |
Soft
hair on outside shell |
Inflorescence |
|
The
first seed |
The
second seed |
Loose
inflorescence.
(
var.) |
Side
inflorescence
(
var.) |
|
Dark
Brown |
Hairiness
|
Hairiness
|
russa
Z.Du |
pilifra
Z.Du |
|
Dark
Brown |
Hairiness
|
Hairlessness
or Hairiness at bottom of awn |
brunnea
Z.Du |
badia
Z.Du |
|
Dark
Brown |
Hairiness
at bottom of awn |
Hairlessness |
hinnulea
Z.Du |
|
|
Grey
|
Hairiness
|
Hairiness |
fumosa
Z.Du |
cineracea
Z.Du |
|
Grey |
Hairiness
|
Hairlessness |
myochroa
Z.Du |
incana
Z.Du |
|
Grey |
Hairiness
at bottom of awn |
Hairiness
at bottom of awn |
airoschistacea
Z.Du |
cieracea
Z.Du |
|
Grey |
Hairlessness
|
Hairlessness
|
elbida
Z.Du |
tephra
Z.Du |
|
Yellow |
Hairiness
|
Hairiness
|
ochracea
Z.Du |
caesariata
Z.Du |
|
Yellow |
Hairiness
|
Hairiness
at bottom of awn |
cerina
Z.Du |
|
|
Yellow |
Hairlessness
|
Hairlessness
|
alutacea
Z.Du |
epipsila
Z.Du |
|
Brown
|
Hairiness
|
Hairiness
|
atrobrunnea
Z.Du |
|
|
Purple |
Red
protect caryopsis |
Chinese |
Sangri |
Sangria
Z.Du |
|
Purple |
Transition
to Nakedness |
Chinese |
Langxian |
Langxiana
Z.Du |
Table 4: Yield of Common wheat,
Durum wheat and Oat in three replicated treatments
|
Treatments
Varieties |
I |
II |
III |
Tt |
X
t |
| Durum
Wheat 281
Rinong
No. 23 (Wheat)
No.
3 (Naked Oat)
Durum
Wheat 291
No.
1 (Naked Oat)
Durum
Wheat 266
No.
2 (Naked Oat)
Zangchun
No. 6 (Wheat)
Durum
Wheat 290
Durum
Wheat 294 |
144.95
219.9
146.6
209.8
236.55
173.25
203.25
116.6
113.3
59.95 |
146.6
173.25
94.95
183.9
266.55
144.95
266.55
134.95
119.95
240.65 |
186.6
203.6
61.65
159.95
199.9
107.95
239.9
146.6
106.6
125.65 |
478.15
596.75
303.2
553.75
703
426.15
709.7
398.15
339.85
426.25 |
159.38
198.92
101.07
184.58
234.33
142.05
236.57
132.72
113.28
142.08 |
| Treatments |
1624.25 |
1772.3 |
1538.4 |
4934.95(T) |
|
| X5 |
162.43 |
177.23 |
153.84 |
|
164.5(x) |
Table
5: Analysis of Variance based on the yield result in table 4. t
| Variation
source |
DF |
SS |
MS |
F |
F0.01 |
F0.01 |
| Treatments
Varieties
Error
margin |
2
9
18 |
2799.94
61043.85
28017.24 |
1399.97
6782.65
1556.51 |
0.899
4.36 |
3.55
2.46 |
6.01
3.60 |
| Total
variation |
29 |
91861.03 |
|
|
|
|
Table
5: T-test at 0.05 and 0.01 levels for the result in the table 5
| Variety |
Average Yield (kg) |
0.05 |
0.01 |
| 2
1
ck2
291
281
274
266
ck1
290
3 |
236.57
198.92
184.58
159.38
142.08
142.03
132.72
113.28
101.01 |
a
a
ab
ab
bc
c
c
c
c
c |
A
A
AB
AB
AB
AB
AB
AB
BC
C |
|