THE INTRODUCTION AND USE OF OAT (AVENA SATIVA) CULTIVARS IN PAKISTAN

 

 

 

 

 

by

Dr. Muhammad Dost

Aga Khan Rural Support Programme (AKRSP), Gilgit, Pakistan

 

 

CONTENTS

SUMMARY

1. INTRODUCTION

2. FARMING SYSTEMS AND CROPPING PATTERNS

3. FODDER OAT INTRODUCTIONS

4. THE ROLE OF OATS

5. ROLE OF FORAGE OATS IN THE NORTHERN AREAS

6. INFORMATION ON FODDER OATS (MAINLY) FROM OTHER COUNTRIES

7. OAT RESEARCH AND INVESTIGATIONS IN PAKISTAN

8. OAT SEED PRODUCTION

9. CONCLUSIONS

10. REFERENCES

SUMMARY

This article describes how the introduction of improved, multi-cut, high yielding cultivars of forage oats (Avena sativa L.) has changed the crop’s status in Pakistan from one limited to a few stations and large farms to one of the most important cool-season fodders in the space of less than twenty years. Livestock, especially cattle and buffaloes, mainly stall-fed, are very important in Pakistan’s agricultural economy. Crop residues, sown fodder, and sometimes a little rough grazing form the basis of the ration with concentrates for commercial dairy stock.. There is a strong and increasing demand for meat and dairy products. Agricultural land is limited so the main way to increase forage availability is through increasing yield per unit area. Much of the agricultural areas have a suitable climate for year-round cropping so green feed, from a range of cold and hot season crops is the main forage supply. Urban dairying is very important and they depend on forage which is grown as a cash-crop for town sale. The introduction of the new oats coincided with a great expansion of the dairy industry which gave an added impetus to commercial forage growing in the irrigated tracts. The main winter fodder had been berseem (Trifolium alexandrinum) which is still grown on a vast scale; it is an excellent forage and yields well in autumn, if sown early and in spring but performs poorly in the coldest weeks of winter which is a major period of forage scarcity; unlike oats, berseem cannot readily be conserved as hay. Berseem is another case of astonishingly successful introduction and uptake by the farming community: it was introduced to Sind from Egypt between eighty and ninety years ago and in twenty years was the main winter forage throughout lowland Pakistan and the northern irrigated tracts of India, displacing the former winter legumes Trifolium resupinatum and Melilotus indica almost totally. The methodology of introduction, screening, selection, field testing and extension are described; along with the very necessary seed bulking and distribution. Special attention is given to the great success of fodder oats in the smallholder areas of the Northern Areas at altitudes from 1,000 m to 2,300 m, where forage is part of the subsistence system, to help over winter stocks which graze alpine pastures when they are free of snow.

World literature on fodder oat cultivation is reviewed, along with national work, in a second section and there is a comprehensive bibliography.

  1. INTRODUCTION

Resolving forage and livestock feed constraints through year round forage availability, as well as the supply of sufficient quantities of milk and milk products, have dominated Pakistan’s agricultural development strategies for almost thirty years.

A chronic fodder shortage, most serious in winter, is a major limiting factor for livestock production. There are two traditional fodder deficit periods, December-January (when the traditional winter fodder crops, especially berseem, the major winter fodder (Trifolium alexandrinum), oats (Avena sativa), shaftal (Trifolium resupinatum), and lucerne (Medicago sativa) are dormant) and May-June (when the main summer season fodder crops such as maize (Zea mays), pearl millet (Pennisetum glaucum), sorghum (Sorghum bicolor), Sorghum-Sudan grass hybrids (Sorghum vulgare var. sudanense) have just begun growth and the winter fodder season is over). Until recently seed of improved fodder cultivars, especially multicut oats have not been available, so there has been a scarcity of fodder both in terms of quantity and quality.

Oats (Avena sativa L. and Avena byzantina C. Koch) rank fifth (Dost, 1997) in terms of world production of cereals. They are also widely used as a companion crop for under-seeding of forage legumes. Oats are mainly grown in temperate and cool sub-tropical environments. In Pakistan, although not used as human food, they are an important winter fodder, in both irrigated and rain fed areas as green feed, hay, silage or grazed.

The average green yield of local cultivars (tall with very narrow leaves and thin stems, hence, not responsive to nitrogenous fertilizers due to lodging) under rain fed conditions is 20 tons/ha (Bhatti et. al., 1992), which is very low and insufficient to provide even maintenance rations for the numbers of livestock kept. In winter farmers have only dried summer grass or dry stalks of summer cereals to supplement the small amount of forage grown and have to purchase costly fodder transported in large quantities from distant, irrigated tracts.

In contrast to local landraces, improved oats grow very fast, can be cut earlier and have considerable potential to provide feed during deficit periods and low temperatures. In Balochistan province, wheat was traditionally used as fodder; however, with the introduction of improved forage oats, use of wheat for fodder has been reduced. Generally, farmers harvest these fodders at 50 percent flowering, or at a later stage to get maximum green yield with a consequent loss in quality (Dost, 1997).

The ideal fodder oat should be high in crude protein (12.42 per cent) and digestibility (in vitro dry matter digestibility - 67.42 percent), and low in crude fibre - acid detergent fibre (33.13 percent). Fodder yield and quality is greatly influenced by plant age, the crude protein content and in-vitro dry matter digestibility decreasing as the forage crop matures. Of course dry matter yield increases with advancing maturity (Dost et al., 1994).

Appreciating the importance of improved oats as a promising source of early, nutritious fodder, and to overcome the winter feed problem, the Government decided to import oat cultivars from countries thought to be the best potential sources of germplasm.

2. FARMING SYSTEMS AND CROPPING PATTERNS

In Pakistan, most of the crop production is in four major agro-ecological regions. With variations in climatic conditions, dependence on irrigation or rainfall, temperatures, soil fertility, land holding size etc., four major farming systems are prevalent:

(i) Medium to high altitude - the mountain/hilly regions such as the northern parts of North West Frontier Province (NWFP), Northern Areas and parts of Balochistan. Those near the Himalaya get reasonable rain; the Gilgit and Northern Area lands are in a rain shadow and rely on irrigation, often spate irrigation deriving from snow or glacier melt. Land holdings are very small and arable areas limited to scattered irrigable alluvial fans. Crops are often undersown in orchards. Agriculture is purely subsistence. Livestock comprise both cattle and small ruminants. Dry stock and small stock may go on transhumance. These areas are 1,000 to 2,300 m above the plains and have cold to extremely cold winters. At medium elevations, maize, rice, potato, wheat, barley, and shaftal are grown; at higher elevations (2000 m), seed potatoes, wheat, buckwheat, foxtail millet, barley, oats, and alfalfa are grown.

(ii) Medium altitude - the rain fed tracts, mainly the Pothowar plateau, Rawalpindi-Islamabad, Chakwal, Jehlum, Attok, Mianwali, D. G. Khan (Punjab), Bannu, Karak, Kohat, parts of D.I. Khan (NWFP), and most of Balochistan have conditions similar to the irrigated areas, but mainly depend on rain for cropping. Large ruminants are important, and forage production is very seasonal; there may be some rough grazing. Major crops are mustard, wheat, barley, oats, and lentils in winter; maize, sorghum, millet, guar, and groundnut in summer. Maize is an important dual purpose crop in high rainfall regions, but it is replaced by groundnut in drier areas.

(iii) The vast irrigated tracts of the plains - Central and Northern Punjab, parts of NWFP, Sindh and Balochistan are where the bulk of Pakistan’s agriculture is, with intensive commercial farming over very large areas. Large ruminants especially dairy buffaloes are important, stall-fed on crop residues and fodder. With warm temperatures and plentiful irrigation, conditions are optimal for luxuriant crop growth for ten months of the year, so year-round forage production is possible. At low altitude, in a monsoon climate, these areas are sub-tropical with searingly hot summers. Wheat, cotton, sugar cane, maize, rice, lucerne, berseem, and oats are other main crops. These areas meet almost all the grain and forage requirements of the urban dairies, including those in rain fed regions.

(iv) Coastal areas - with sub-tropical conditions, such as south Sindh and parts of Balochistan. Pearl millet, sorghum, maize, sugarcane, barley, and oats are common. The rainfall is scattered and erratic and irrigation is a factor of extreme importance, allowing cultivation in difficult areas. Amongst climatic factors, temperature and water availability are the major ones. Soils with regards to both their physical and chemical properties, which influence water holding capacity and fertility, affect production systems. These factors also influence pH and salinity levels of the soils. The major winter forages are oats (Avena sativa), berseem (Trifolium alexandrinium), lucerne (Medicago sativa), vetch (Vicia villosa var. varia), barley (Hordeum vulgare), mustard (Brassica spp.); the summer forage crops are maize (Zea mays), sorghum (Sorghum bicolor), sorghum sudan grass hybrid (Sorghum vulgare x. sudanense), pearl millet (Pennisetum glaucum), cowpeas (Vigna unguiculata), and guar (Cymopsis tetragonoloba).

In most rain fed regions, farmers grow local landraces of traditional forages such as sorghum, millet, mustard, and barley with inherent low yield potential and poor nutritive value. Livestock are fed on dried maize-sorghum-millet stalks and graze wild grass of extremely low nutritive value. In winter, wheat is cut and fed. Poor nutrition results in poor health and production. In prolonged droughts, even productive animals are sold to get cash for domestic needs. In the rain fed tracts, forage availability is the major criterion in deciding which kind of stock to keep.

Wheat and maize in rain fed areas are subsistence crops; groundnut and melons are cash crops. Sorghum with maize, and mustard intercropped in wheat, are the major green forages, supplemented by maize thinning and weeds. Barley and oats are special forages generally grown for sale near big cities and urban areas, mainly under tubewell irrigation. Several thousand tons of oat forage are transported to urban dairies daily in season. Forage fetches reasonable prices in winter for the needy farmers. Thousands of urban dairies, generally without land for forage production, rely on daily purchase. In dry seasons, these peri-urban commercial farms cannot meet the forage requirements of city dairies. Due to the large gap between forage supply and demand, it is transported from irrigated areas hundreds of kilometres from some big cities. In rain fed areas wheat straw and sorghum/maize/ millet dry stalks are the bulk of feed. In addition to green and dry forages concentrates are fed to animals in milk.

Livestock rearing has evolved specialized crop management practices e.g. intercropping of companion crops, maize thinning and fallowing of land help to feed animals. Farmers in rain fed areas have developed cropping patterns in response to rainfall, moisture, manure availability, soil fertility, and forage requirements. Of all these factors rainfall is the most important. There are therefore, three very different fodder situations:

(i) First, in rain fed areas agriculture is for subsistence and crops depend on rainfall. Forage requirements are partly met through forage crops and partly by grazing fallows;

(ii) Second, in the irrigated tracts peri-urban forage production is mainly for commercial purposes, and

(iii) Thirdly, in high altitude mountain regions both forage production and stock rearing are for subsistence and forage requirements are partly met through cultivated forage on very small holdings where irrigation is available and partly through using alpine grazing to the maximum extent.

On the plains there are two very different types of stock rearing: subsistence and commercial; the latter in the peri-urban or "milk shed" areas of towns, with some even in town. There are also two levels of fodder production; some farmers grow for their own stock (again sub-divided for subsistence and commercial) while others grow fodder as a cash crop and may not have livestock. Commercial forage farmers practice very intensive cultivation and usually produce four crops per year from the same piece of land due to optimum temperatures, availability of irrigation, proper planting time and management, with application of balanced doses of fertilizers with different times of application to sustain soil fertility for maximum/early crop growth and productivity.

Commercial dairies are rare in the rural tracts but very important around big cities such as Karachi, Hyderabad, Lahore, Sheikhupura, Gujrat, Gujranwala, Rawalpindi-Islamabad, Faisalabad, Peshawar, Charsada, Nowshera, Quetta, and Mirpur, AJK. Feed for these dairies is transported by truck, tractor trailer, and camel/bull/donkey carts for forage produced nearby, but some is produced and transported from over 300-400 km away e.g., from Hyderabad/ Sukkar/Nawab Shah to Karachi; from Kasur, Sheikhupura, Gujranwala, Faisalabad, and Renala Khurd to Mirpur (AJK) and Rawalpindi-Islamabad, and from Nowshera, Charsada, Mardan, and Malakand to Peshawar. Cities in irrigated regions have relatively close forage sources, but Karachi, Islamabad-Rawalpindi, and Mirpur (AJK) must purchase feed from distances of 300-400 km. Buffaloes predominate in town dairies of Punjab, NWFP, and Sindh, as their milk is preferred to cow milk and fetches a higher price, essentially because of its higher fat content (8.5 vs. 4.5%). In Quetta, Balochistan, less than half of the urban herd is buffalo, due to several factors including lack of irrigation, scarcity of green and dry forage and because buffaloes require nearly twice as much forage and feed as cows. Afghan refugees brought their pure and cross Friesian cattle to Quetta with them and their excellent condition and performance has attracted local dairymen to cattle.

Rising transport costs have increased the making of oat hay. Data are not available, but huge quantities of oats are now regularly made into excellent hay, baled and transported over long distances. In the 1950s and 1960s, oats were grown for fodder, but mainly to make hay for horses, then the main transport animal in rural and urban areas.

The rapid rise in the popularity of oats over the last fifteen years has been due to the introduction of high yielding cultivars that provided several cuts, such as cvs Scott, S-81, Tibour, Cuscade, Swan, and PD2-LV65. In the late 1980s the dairy industry began to expand and more and more milk marketing outlets became available. A large number of milk processing units have been installed. The expanding urban population has provided a lucrative milk market and dairy farming is now better managed.

Forage production is an important business especially near big cities; a range of crops are grown to maintain a year-round supply. Improved, multi-cut oats are very popular in urban irrigated areas such as Kasur, Sheikhupura, Gujranwala, Faisalabad, and Renala Khurd (Punjab); Nowshera, Charsada, Mardan, and Peshawar (NWFP); and Hyderabad, Sukkar, and Nawabshah in Sindh. Oats have almost replaced the poor quality wheat and rice straw that was the basis of winter feed; berseem provides very high-quality cool season forage and is marketed in vast quantities, but production tends to peak in spring and is poor in the coldest months – it cannot easily be made into hay. Improved oats provide forage in cold weather when other green feed is scarce and are replacing the forage brassica which were formerly used in the winter gap (Suttie, 2000).

Nevertheless, forage yields are low compared to their potential. Improved cultivars and technology have been slow to reach the small scale farms which account for the bulk of forage production. Also seed production has lagged behind plant breeding and introductions. Recent medium scale on-farm work has indicated that yields can be raised two to three times with the use of available improved varieties and appropriate agronomic techniques. In an area where land and irrigation are the major limiting factors to enhancing fodder production, intensification is the only way to meet the country’s needs for livestock products and hence, forage.

3. FODDER OAT INTRODUCTIONS

Fodder oats were introduced during the early British era, but it was only in the 1970s that 400 oats cultivars were acquired from Australia, Canada, Europe, New Zealand and USA. These included materials donated by NZ DSIR/NZ CFRI in 1979 and form the basis of Pakistan’s fodder oat improvement programme. Further commercial importations were made under the World Bank (Hill Farming Project) for Azad Kashmir in the 1980s. Some cultivars from this material still play an important role in the provision of green-feed or hay across a wide range of ecologies. The Fodder Research Programme, National Agricultural Research Centre (NARC), which has the national mandate in fodder improvement, also introduced several improved oat cultivars from Western countries in the mid 1980s. During the late 1980s, the FAO Afghan Programme arranged for fodder oat material to be multiplied with private seed companies in Pakistan, and under Government supervision at research centres.

[See the Annual Report of the Pakistan Agricultural Research Council (PARC) which includes the work of the National Agricultural Research Centre (NARC), Islamabad.]

The Fodder Research Institute, Sargodha , is the only research institute in the country to handle the need for seeds of improved or promising forages. It used to be a small station and Military Farm in the suburbs of Sargodha producing fodder for horses, mules, and milking animals for military needs. The Research Institute has 200 hectares of irrigated land for breeding, evaluation, screening, management, selection, and recommendation of forages suited to various agro-ecological conditions in general and especially for the vast irrigated areas of the country. Since the Institute has a relatively large area of irrigated land, seeds of almost all the improved or promising forage cultivars are multiplied.

The Institute has sub-stations in Faisalabad and Bahawalpur for screening under different climatic conditions. Due to easy availability and access of the farming communities to seeds of improved forages, especially oats in Sargodha and Faisalabad, both cities have now become a major source of fodder as a cash crop which is transported and traded in large quantities and hauled over great distances throughout Punjab. The forage cultivars (oats and others) released by the Fodder Research Institute, Sargodha and other research stations are listed below:

Forage crops cultivars released by various Institutes in Pakistan

Crop/cultvar Institute/Station Year of Release

(i) Oats

(ii) Barley

(iii) Berseem

(iv) Maize

(MaizexTeosinte)

(v) Millet

(vi) Sorghum

(vii) Sorghum-Sudan grass Hybrid

______________________________________________________________________

The Fodder Research Institute(FRI) Sargodha CCRI which works under the Ayyub Agricultural Research Institute, Pirsabak, NWFP (Cereal Crops Research Institute, Pirsabak, NWFP), AARI, Faisalabad (Ayyub Agricultural Research Institute, Faisalabad) MMRI, Sahiwal (Maize & Millet Research Institute, Sahiwal), RARI, Bahawalpur (Rainfed Areas Research Institute, Sahiwal).

Improved oats from Sargodha, other research centres and private seed companies have been grown and used routinely by farmers, usually around big cities (from where fodder is transported daily for sale to private and government commercial dairy units and army dairy farms) in lowland rain fed areas, as well as in mountain areas with extremely limited land holdings and irrigation facilities, for some 20 years. In October-November the summer fodders are finished and most winter forages are not yet ready for harvest so there is an acute shortage of fodder. Oats provide assured feed early in the winter if sown at the end of August or beginning of September to be cut at the end of November - beginning of December.

Oat Seed Multiplication and Extension:

The Fodder Research Programme, National Agricultural Research Centre (NARC), Islamabad is responsible for introducing the seed of almost all the new varieties of various forage crops in Pakistan. All the introduced forage crop varieties are evaluated for forage and grain yield traits at NARC, Islamabad.

In the meanwhile, seed is multiplied for further evaluation. Based on the initial yield evaluation and performance data, some promising introductions are selected for evaluation and multiplication in different agro-ecological zones. The Fodder Programme NARC has forage evaluation and multiplication cooperative substations in the four provinces of the country. The selected cultivars are further evaluated in national uniform evaluation trials at all substations under the supervision of scientists from the NARC, Islamabad. As most of the forage research substations have sufficient area for seed multiplication, those cultivars suited to their agro-ecological region are initially multiplied at these stations. Also, the Fodder Programme, NARC, Islamabad and Fodder Research Institute Sargodha have larger areas for seed bulking; therefore, the maximum quantities of all the selected oats are multiplied by both programmes. The seed thus bulked is sold to small farmers, commercial growers, both Government and private dairy farms and other agencies/organizations interested in improved oat fodder production and development.

In Pakistan, most of the seed of improved forage and grain crops is usually multiplied by private seed companies, Government Research and Extension Centres and to a limited extent by commercial growers. The seed thus multiplied is sold to the interested growers throughout the country. The bulk of the improved seed is mainly purchased for seed multiplication in the irrigated regions and the surplus quantities of seed is sold to the private dealers who transport the seed for sale in rural areas. These private local seed dealers are the main source of supply of improved oat seeds to the small as well as big commercial farmers.

In the near past, no small farmer knew about the potential of forage oats cultivars for livestock and never grew oats. Under the Productivity Enhancement Project in 1997-98, large scale demonstrations and seed multiplication of improved oats was done on farmers’ fields throughout the country. Village seed producers were linked with the private seed companies and private seed dealers in all cities for sale and purchase of improved oats seed. The seed companies and seed dealers became a major source of future oats seed bulking and so improved its availability to the small farmers and seed producers. The seed procured by the seed and private dealers in almost all the big towns is now regularly bought and sold in local markets in the rural areas. Most farmers save a lot of their own seed from the initial supply from the seed merchants.

Also, the non government organizations such as National Rural Support programme (NRSP), Sarhad Rural Support Programme (SRSP), Punjab Rural Support Programme (PRSP), and Balochistan Rural Support Programme (BRSP) are working at grass root level in rural areas over almost 70 % of the country. These rural support programmes played a very important role in demonstrating improved forage crops, especially oats. Interested communities indicate their needs for seeds of oats and other improved crops. The representatives of rural support programmes arrange procurement from reputable sources and deliver it to the farmers homes. In this way seed of improved oats became accessible to the small farmers at the village level. This has helped in the quick dissemination of improved oats seeds, even in the remote areas throughout the country.

For the last several years, the Fodder Programme NARC, and private seed companies have been actively involved in the seed multiplication of improved oats for the FAO Afghan Programme for use in Afghanistan.

4. THE ROLE OF OATS

Forage oats (Avena sativa L.) are grown throughout Pakistan in winter under a wide variety of soil and climatic conditions. Improved oats cultivars are now the main source of winter and spring forage in the plain irrigated / rain fed and high altitude (1000-2300 m asl), colder regions, especially in the Northern Areas. They are grown on more than 35 percent of the available arable land under forage crops throughout Pakistan. Grain yields vary from 2.4 to 3.2 tons/ha in the better environments, but less at higher altitudes (2,300 m). In Northern Areas at an altitude of 2,000-2,300 m, the highest recorded green fodder yield ranged from 80-120 tons /ha.

It is an especially important crop in the plains during the cooler autumn, winter and spring months and in the mountains during autumn, spring and early summer. Reported green fodder and dry matter yields are generally higher in some parts of the North West Frontier Province (NWFP) and Balochistan provinces than in the lowlands and mid hills (up to three times higher through the use of locally bred or selected cultivars of western origin), due mainly to better inherent soil fertility combined with the widespread use of chemical fertilizers, farm yard manure and higher incoming radiation (less fog and cloud cover). In the Northern Areas, oats have proved to be invaluable for feeding milking animals especially the stall-fed cattle near or around villages and the bigger cities (such as Gilgit and Chilas, Skardu to meet domestic milk needs), and hence, not able to use traditional grazing lands.

Temperate and cool sub-tropical conditions are congenial for the growth of oats. A well distributed rainfall of 400 mm and an optimum temperature range of 16-320 C from September to April are sufficient to meet its requirement as a fodder crop. Oats can provide green fodder after 60-70 days in an emergency to tide over the scarcity period, but in 90-100 days after germination, large quantities of fodder are produced. Oats are mostly fed green, but any surplus is made into hay. It is a favourite feed of all animals and the straw is soft and much superior to wheat and barley. It is high in total digestible nutrients (TDN), digestible crude protein, fat, vitamin B1 and minerals such as phosphorus and iron. The oat grain is a particularly valuable feed for horses, dairy cows, poultry and young breeding animals of all kinds.

Improved oats have been especially successful for stall-feeding across a wide range of altitudes, climates and ecologies from 400 m to higher than 2,300 m. In the plains fodder oats are grown for dairy cows and buffaloes kept in and near big cities to meet the considerable milk demand; so much of the area under fodder oats is near large cities, and on military and private dairy farms. Forage (including crops other than oats) is transported daily to the cities and sold to feed milch cattle and horses kept for urban haulage.

5. FORAGE OATS IN THE NORTHERN AREAS

New oat cultivars were first introduced to the Northern Areas by the fodder component of FAO Project PAK/86/027, Gilgit. They have proved to be valuable in the Northern Areas, since they grow much earlier and more vigorously than the traditional winter cereals. Green oats is cut in Gilgit and Chilas when no other green feed is available. Higher yields (up to three times) are produced in the 1,000 to 2,300 m band compared to lower altitude zones, possibly reflecting better agro-ecological adaptation. Many cultivars have been positively evaluated at different altitudes in winter in lowland areas through to high alpine areas around 2,300 m.

Feeding of green oats to cows in winter increased milk yields from 1 to 4 litres per day. Not all milk is sold; additional production increases family dietary quality over traditional practices, especially important for the young and aged. (Milk and dairy products are valued constituents in Pakistani diets). Fodder oats, vetch, lucerne, shaftal, and berseem seed are produced for further cultivation, barter and sale to earn extra cash for household requirements.

6. INFORMATION ON FODDER OATS (MAINLY) FROM OTHER COUNTRIES

6.1. Characterization of a dual-purpose crop:

Hadjichristodoulou (1983) described a dual-purpose crop as one suited to use as "grazing plus hay" or "grazing plus grain", Yau and Mekni (1985) classified dual purpose barley cultivars (see Figure 1) as having high forage yield at grazing time and high grain yield after grazing. This was considered a "narrow-sense" definition; in the "broad sense", dual purpose barley should perform well whether grazed or not. Therefore both forage and grain material should be used as resource material in breeding dual-purpose cultivars. In Cyprus, Hadjichristodoulou (1983) concluded that high-grain-yielding lines could be used as basic material for selecting dual-purpose varieties. By contrast, Yau and Mekni (1985) reported that improved high grain yielding barley genotypes generally did not perform well when subjected to grazing. Anderson (1985), however, found no clear differences between dual purpose and forage cultivars in their response to grazing over years as reflected in dry matter and grain yield.

Grain yield after clipping grain type

 

dual-purpose type

 

forage type

Dry matter yield at tillering

Figure 1. Classification of barley genotypes according to their dry matter-yield at the tillering stage and grain yield after clipping (after Yau and Mekni, 1985).

6.2. Desirable features in a dual – purpose cereal crop:

A successful dual-purpose crop should provide a substantial amount of forage and recover quickly from defoliation so that a good yield of grain can be obtained. From the literature, it would appear that such a crop for southern Australia should have the following features:

- Semi-prostrate or prostrate growth habit

- Long pre-jointing phase

- Free-tillering habit

- Leaf canopy of relatively prostrate foliage in the early, pre-jointing phase, becoming more erect as stem elongation occurs and new leaves are higher in the canopy.

- Late flowering time i.e. long vegetative phase, flowering time controlled by a relatively strong response to vernalization. A photoperiod response may also be necessary with early sowing of even a strongly vernalizing type to ensure flowering after risk is passed.

These characteristics would allow early sowing, as soon as moisture is available, to take advantage of favourable conditions for photosynthesis in the autumn. Substantial herbage production should be achieved in the early stages of growth and could be safely grazed before flower initiation and stem elongation. If defoliation is not too severe, the type of plant envisaged should retain some leaf after grazing, and should be able to rebuild a canopy and re-tiller as necessary, so that when stem elongation occurs, an adequate number of fertile tillers should be present to ensure a good yield of grain. Because of late flower initiation due to the vernalization requirement, frost damage should be avoided, whilst with only a relatively weak response to photoperiod, flowering and grain production should not be delayed sufficiently for summer moisture stress to be a problem. Whether more than one grazing would be possible would depend on the degree to which various characteristics were expressed in particular cultivars and on environmental conditions and grazing management.

6.3 Forage Quality:

Cattle can be maintained in good condition on oats at a time of the year when the supply of other feed is scare and costly. Many cultivars provide good forage if cut at flowering or soon after. Broad-leaved cultivars produce a higher forage yield, but narrow leaved ones are preferred by horses and cattle. The demand for meat and dairy products is increasing due to the rapid population growth in Pakistan, so improvement of livestock production is urgently needed and quality forage plays a pivotal role. Although 16-19 % of the total cropped area in Pakistan is planted to fodder, animals are generally underfed. To operate an efficient and economical livestock industry, high yielding, nutritious and multi-cut fodder oats are needed to feed more animals (Dost, 1997).

Hussain et al., (1993) reported that ‘Fatua’ oats harvested at various intervals produced more fodder and less crude protein with plant age/advance in crop maturity. The crop should be harvested at a stage that provides an optimum compromise between forage yield and quality. Maximum green fodder and dry matter yields and crude protein contents were recorded when oats was harvested at 50 percent flowering. Hussain et al. (1994) also reported that the highest green and of course dry matter yields of five oat cultivars were at 50 percent heading.

Dost et al. (1994) concluded that forage yield, dry matter yield, and crude fibre increased while seed yield and crude protein declined with advancing maturity. Harvest at 50 percent flowering resulted in superior forage and dry matter yields with inferior nutritive forage value as determined by lower crude protein and higher crude fibre contents, compared to harvesting at the vegetative stage at 70 and 85 days after planting. Minimum forage and dry matter yields with maximum forage quality resulted from harvesting at the mid vegetative stage at 70 and 85 days after planting.Young cereal plants provide excellent quality herbage which is highly nutritious for lactating ewes and young growing lambs. Their digestibility, crude protein content, carotene, mineral and vitamin contents have been reported to decline with plant age, whereas crude fibre and nitrogen free extract increase (Skorda, 1977; Eagles et al., 1979; McDonald and Wilson, 1980). However, yield per unit area of both crude protein and digestible dry matter increase with advancing maturity until the milky stage (Hadjichristodoulou, 1976a & b; Hughes and Haslemore, 1984; Droushiotis and Wilman, 1987).

High quality feed, suitable for growth and lactation, should exceed values of about 67 percent in digestibility and 13 –15 percent in crude protein; less than 65 percent digestibility indicates feed suitable for maintenance only (Raymond, 1969; Hughes and Haslemore, 1984). Whilst Hughes and Haslemore (1984) concluded that if the digestibility of a forage is satisfactory (above 67.5 percent), the protein content will generally also be satisfactory (above 12-15 percent), Raymond (1969) and Eagles et al. (1979) found that decline in digestibility of forage crops (including cereals) led to a reduction in voluntary intake by animals.

Several workers (Lassiter et al. 1958; Zogg et al. 1961) report comparisons of maize and oat silage for milking cows. Both concluded that maize silage was superior to oat silage for milk production. However, in each study the silages were not fed alone and a considerable portion of the dry matter intake was from hay.

Mowat and Slumskie (1971) indicated that maize silage contained more digestible energy than barley silage when fed to finishing steers. The low digestible energy content of barley silage was also demonstrated by Fisher et al. (1972) in a study with dairy cows. Neither report assessed the suitability of the crops in terms of both field yields and feeding value.

Under Tasmanian conditions, Abdul-Rahman et al. (1985) found that very early-sown (4 January) oats still had a digestibility of 60 percent and crude protein content of 14 percent in June, when the crop was close to flowering. It is noteworthy that under the long growing season of high rainfall areas, crops always produce high dry matter digestibility (75-85 percent) in winter (Dann et al., 1977; Eagles et al., 1979). The relationship between digestibility, radiation and temperature has not yet been shown experimentally, but sunny days followed by cool or cold nights should give high levels of soluble carbohydrates (Wheeler, 1981). Whilst lower digestibility values were reported in low rainfall areas, crude protein content at later stages of crop development, by contrast, was higher (Skorda, 1977; Droushiotis and Wilman, 1987). This was attributed to less leaching of soil nitrogen in low rainfall areas (Hadjichristodoulou, 1976).

6.4 Maturity period:

Although grazing and cutting contribute to delay in flowering, a late flowering characteristic is important in dual-purpose cereals, in many places in high rainfall areas, to avoid the risk of frost damage . Through the findings of many researchers, time from sowing to heading or maturity is a good indicator for the selection of dual-purpose crops. In the high rainfall areas of Punjab, the recommended dual-purpose oat cultivar S-81 was the earliest to mature after heavy grazing/cutting. It was, however, the latest when ungrazed and grown for grain (Dost, 1994). This trait of late maturity in a dual-purpose crop was confirmed by McLeod et al, (1985) with a cutting height in oats of 2 cm. Hadjichristodoulou (1983) and Yau et al., (1987) concluded that dual-purpose lines tended to be later heading than the usual grain types. Furthermore, the time to heading or maturity appeared to be the main factor affecting yields, while tiller number, head number and plant height were less important (Yau et al., 1987). But these authors warned that, although late lines had better recovery to produce high grain and straw yields after simulated grazing, they were not good dual-purpose types since they gave little at the time of defoliation.

6.5 Factors affecting the response to date of sowing:

The response to early sowing is apparently a valuable criterion of a dual-purpose crop. This advantage in high rainfall areas in Australia may be associated with a response to vernalization (Anon, 1986; Dann et al., 1977; Davidson et al., 1985a), which gives flexibility in sowing time with stability in flowering time. Varieties responding to long days (Davidson et al., 1985b), may show a similar delay in flowering and hence escape damage by water logging and frost.

Johnson and Dann (1984) suggested that further improvement for dual purpose use could come from breeding cultivars with a higher vernalization requirement, permitting flexibility in sowing dates from late summer to mid autumn without risk of frost damage.and it is very important in areas with prolonged freezing temperature especially in the high altitude regions. For many cereal growing areas, however, these cultivars should also be able to flower early in spring (little requirement for long photoperiod), so that grain is produced before the onset of early summer moisture stress.

Stern and Kirby (1979) stated that vernalization, photoperiod and temperature are the most important environmental stimuli on phenological events as they influence tillering, spikelet initiation and development of cereals. It is difficult to separate the effect of these factors on phenological events in cereals.

6.6. Effect of nitrogen fertilization on dry matter yield:

Many workers in Australia have found that nitrogen fertilization at various levels (ranging from 67 to 180 kg N/ha) increased the forage yield of oats and that this was a good method of overcoming winter feed shortages (Archer, 1969; Blunt and Fisher, 1976; Crofts, 1966a; Wheeler, 1968). A better response to nitrogen was achieved in spring when the temperature rose (Cook and Lovett, 1974).

Leaf area index (L), total living dry weight, stem dry weight and number of tillers were all significantly increased by nitrogen fertilizing of defoliated and non-defoliated oats (Cook, 1971). Mehra et al. (1971) found that fodder yield was positively correlated with plant height, leaf length and leaf width, tiller number, and stem girth.

The number of tillers at the end of stem elongation and the final number of tillers with ears in cereals were reported to be increased by the addition of nitrogen, especially when applied early, thereby increasing yield (Needham and Boyd, 1976; Graham et al., 1983; Garcia del Moral et al., 1984). A nitrogen deficit reduces tillering due to:

- Retarded appearance of lateral buds (Hewitt, 1963, cited by Garcia del Moral et al., 1984).

- Limited root growth (Briggs, 1978; Cook, 1971).

- Small weak shoots whose leaves contain reduced levels of chlorophyll and carotenoids (Briggs, 1978).

By contrast, however, some workers have demonstrated that nitrogen fertilizer neither increased dry matter yield significantly nor compensated for the deleterious effect of clipping on the grain yield of oats and barley. This might be due to one or both of the following reasons:

- Available soil nitrogen was already high (Dann, 1971), where the site had previously been under leguminous pasture (Spurway, et al., 1976), or a heavily manured crop (Gardner and Wiggans, 1960) or fallow (Brown, 1975).

- Nitrogen fertilizer was applied in early winter when the temperature was very low (Archer, 1969). Cook and Lovett (1974) suggested that the ambient temperature rather than nitrogen supply is the limiting factor in growth of oats during winter.

7. OAT RESEARCH AND INVESTIGATIONS IN PAKISTAN

The Co-ordinated Fodder Research Programme NARC, Islamabad vigorously evaluated 400 introduced cultivars (1970s introductions) throughout Pakistan and selected 20 promising ones based on maximum forage yield, dry matter yield, maturity etc. The 20 selected cultivars were further evaluated in all four provinces in the autumn (rabi) season under a wide variety of soil and climatic conditions to select and recommend the most promising and suitable ones for different areas within diverse agro-ecological regions. The variation in environmental conditions in different agro-ecological regions is well known to plant breeders. Fluctuations in rainfall and temperature are neither consistent nor predictable from year to year or from place to place. Fertility status and soil types also vary throughout the country. Serious losses in forage yields due to drought, disease, and insects are common in many areas.

Pakistan has a long history of selecting and using local oat landraces, and in modern times, especially the last twenty years, breeding and testing of improved cultivars incorporating material from western sources into local fodder programmes has been a continuous process. Year-round fodder production is the most important component in the farming systems and a great deal of research is being carried out in the four provinces of Pakistan as part of a national coordinated fodder research programme. The details are presented below.

7.1 Genotype x Environment Interaction and Screening Methods:

Considerable genotype by environment interaction has been noted across latitude, altitude, seasonal sequence, with some cultivars producing significantly better forage yields than others in certain environments and management regimes. So far, this is poorly documented and has only been modestly exploited in Pakistan due to limited local resources. Effectiveness of varietal testing programme is influenced by several factors. These include experimental design, the number of locations, and the number of years used to average variety means. Information is required as to whether forage oat varieties respond differently when planted under diverse environment interactions, and if so, how important such genotype x environment interactions might be in an oat variety evaluation and selection programme.

Horner and Frey (1957) report that dividing the Iowa State into 2, 3, 4 and 5 sub-areas would reduce variety x location interaction effects by 11, 21, 30 and 40 percent respectively and hence increase the efficiencies of the state wide oat performance trials.

Liang et al. (1966) studied genotype x environment interaction in wheat, barley, and oats and recommended dividing Kansas into 3 and 4 areas, for barley and wheat testing, respectively. It was concluded that it was not necessary to divide the state into smaller areas for oat testing.

Dost et. al. (1993) studied the optimum allocation of resources in varietal evaluation for 13 oat genotypes for forage yield at 4 locations for 4 years and suggested that the three provinces under study should be divided into sub-areas on the basis of variation in rainfall, temperature, soil type, and soil fertility in order to minimize genotype x environment interaction for an efficient forage oat evaluation programme (Table 1).

Table 1. Expected Variance of a Variety Means (Vx) for various assumed Numbers of Replicates and Locations per Test.

No. of replicates

No. of years

2

4

No. of Locations

No. of Locations

2 4 6 8 10 12 2 4 6 8 10 12
2 2.25 1.29 0.97 0.81 0.71 0.65 1.18 0.67 0.50 0.42 0.37 0.34
3 2.22 1.27 0.96 0.80 0.70 0.64 1.16 0.67 0.50 0.42 0.37 0.34
4 2.20 1.26 0.96 0.79 0.70 0.64 1.16 0.66 0.50 0.42 0.37 0.33
5 2.19 1.26 0.95 0.79 0.70 0.64 1.15 0.66 0.50 0.42 0.36 0.33
6 2.18 1.26 0.95 0.79 0.70 0.64 1.15 0.66 0.50 0.41 0.36 0.33

Dost et al. (1993)

To evaluate the importance of genotype x environment interactions, De Pauw et al. (1981) compared three cultivars of wheat, three of oats, and three of barley for four years at five locations in north-western Canada. They concluded that cultivars showed differential responses in specific environments, which can be used to determine areas of cultivar adaptation.

In a study on genotype x environment interaction at twenty-seven locations for five years, in Alberta, Canada, Kibite et al. (1988) suggested the need to divide Alberta into six rainfed and two irrigated areas.

7.2 Oats as a multi-cut crop:

Compared with wheat and barley, the traditional sources of arable green winter fodder in Pakistan, oats provide multiple cuts, tiller profusely and yield more, and are of higher nutritional value. A major reason for this is that the standing oat crop can be harvested progressively, releasing land earlier than normal for follow-up crops or relay cropping. Late in the season, any remaining crop can be cut at the farmer’s discretion and dried as hay. This coincides with optimum soil moisture for land cultivation and planting of the following crop. This also allows small areas or peripheral lines on terraces to be saved for seed production. In many, but not all instances, more recently bred cultivars out-yield older ones (Dost et al. 1994).

Although the utilization of oats as a multi-cut crop is a common practice in Pakistan, relatively little research data are available to aid the farmer in the selection of the best forage harvest schedule for the dual utilization of oats to allow one to obtain high forage yield, grain yield, and good forage quality.

Bhatti et al. (1992) evaluated 13 oat cultivars under a two cut system at NARC, Islamabad during 1985-86 and 1986-87. The results of the study revealed that the cultivars PD2LV65 and S-81 produced 28.05 percent and 26.24 percent more green forage yield and 26.30 percent and 21.93 percent more dry matter yield respectively in two cuttings compared with the control variety. Thus oat cultivars PD2LV65 and S-81 were found suitable for multi-cut systems both under irrigated and rainfed conditions (Table 2).

Table 2. Green and dry matter yield of various oat cultivars at the National Agricultural

Research Centre, Islamabad during 1985-87 (average of two years)

Cultivar First Cutting Green fodder yield 2nd cutting Total Percentage increase (+) decrease (-) over control First cutting Dry Matter yield 2nd cutting Total Percentage increase (+) or decrease (-) over control
‘DN-8’ 27.15 ab 45.99c 73.14 + 7.22 5.185 17.936 23.121 + 6.31
‘Algerian’ (control) 25.46 ab 42.75cd 68.21 - 4.862 16.886 21.848 -
‘W. No. 11’ 23.30b 42.28cd 65.68 - 3.85 3.984 16.892 20.876 - 4.01
‘Avon’ 23.76b 45.21cd 68.97 + 1.11 3.944 17.632 21.576 - 0.79
‘Fulgrain’ 27.47ab 44.75cd 72.22 + 5.88 5.246 17.452 22.698 + 4.37
‘Sargodha-81’ 29.32a 56.78a 86.10 + 26.24 5.893 21.576 27.469 + 26.30
‘Golden rein’ 29.16a 43.67cd 72.83 + 6.78 5.103 15.502 20.605 - 5.25
‘Swan’ 26.85ab 49.84b 76.69 + 12.43 5.235 17.792 23.024 + 5.86
‘PD2-LV65’ 29.32a 58.02a 87.34 + 28.05 5.805 20.713 26.518 + 21.93
‘Kent’ 28.40a 45.37cd 73.77 + 8.15 4.629 16.424 21.053 - 3.19
‘Java Lahori’ 27.77ab 41.51d 69.28 + 1.56 4.804 15.151 19.955 - 8.24
‘A. fatua’ 26.85ab 45.37cd 72.22 + 5.88 4.994 16.877 21.871 + 0.56
‘Ealge No. 1 25.00ab 44.14cd 69.14 + 1.36 4.550 16.552 21.102 - 2.97

Means followed by the same letters do not differ significantly at 5 % level of probability.

Bhatti et al. (1992)

7.3. Oats as an intercrop/companion crop:

In order to obtain early and good yields under small land holdings and severe winter conditions, compatible fodder crops of different types might be planted in mixture to produce high fodder yields and forage quality per unit area per season (Table 3). The leguminous dwarf fodders like berseem/vetch can be mixed with tall growing fodder like oats, rye grass, brassica etc.

Oat + vetch and barley + vetch combinations produced on average 110 and 70 tons/ha of green material compared with 100 and 56 tons/ha pure oat and barley stands respectively within the 1400 to 2000 m altitude band (Table 3).

Table 3. Green and dry matter yields (tons/ha) of

oats, barley and vetch during 1994-1997

Cultivars

Sites

Gilgit Chilas
Green yield Dry yield Green yield Dry yield
Oats 100 21 105 23
Oats + vetch 110 24 102 22
Barley 56 12 59 14
Barley + vetch 70 16 76 18

Also a deep-rooted crop like Lucerne can be mixed with shallow rooted crops like oats, rye, barley or brassica. The annual fodder is usually planted in the space between the rows of perennial odder. This technology has been practically demonstrated to the farmers of the Northern Areas. There are many advantages of mixed planting over sole planting:

- More than one crop per season per unit area

- Weed control is easier

- More fodder yield and better quality of fodder

- Maintains and improves soil fertility.

Oats were inter-cropped in winter active Lucerne and red clover planted in rows spaced at 30 cm apart at a number of locations. The mixture of Lucerne + oats, red clover +oats, and berseem + oats produced maximum green and dry matter yields as compared to the sole crops of either legume. The results are presented in Table 4.

Using oat + berseem mixtures provided early and more fodder yields, increased milk production by 20 litres per animal per month on average, compared with traditional practices. At the same time, the demand for purchased concentrates was reduced by 20 kg/month per animal and milk production was extended by an extra two months (Table 4).

Table 4. Green and dry matter yields (tons/ha) of various legumes

and oats cultivars at Gilgit during 1993-1994

Treatments Forage yields Dry matter yields
Lucerne alone 70 18
Lucerne + Oats 115 30
Berseem 80 17
Berseem + oats 135 30
Red clover 63 16
Red clover + oats 94 26

To overcome the problems of fodder shortages due to limited land holdings and also to obtain maximum fodder per unit area per season, multiple cropping/ mixed planting techniques were tried. The details are presented below.

Non-winter-dormant Lucerne cultivars (Sundar being the main one), introduced by the FAO project PAK/86/027, Gilgit, have been very successful (Table 5). In some cases they may suffer some frost damage at high altitudes, but they grow throughout the year and yield more than twice as much as the landraces in the low altitude double crop areas 2000 m and below. The new cultivars also provided maximum green feed in the critical December-January period when traditional crops are usually dormant and the area experiences a fodder deficit period.

Table 5. Effect of mixed planting on green and dry matter yields (tons/ha) of lucerne, red clover, and oats in 1996-97

Treatments

Sites

Sultandabad

Rahimabad

Saling

Green yield Dry yield Green yield Dry yield Green yield Dry yield
Lucerne Sundar

110

30

105

26

68

20

Red clover

60

16

62

18

50

13

Oats

100

32

95

30

80

26

Average

90

26

87.33

24.66

66

19.67

Lucerne + oats

140

39

136

37

102

30

Red clover + Lucerne

115

32

108

26

70

22

Red clover + oats

90

26

93

28

75

23

Average

115

32.33

112.33

30.33

82.33

25

Pendleton (1957) reported that oats seeded at a low rate in 32 inch rows as a sole crop returned 190 bushels for each bushel seeded, compared to the 42 bushel return from a regular seeded 8 inch row spacing inter-seeded with clover. Also red clover inter-seeding reduced spring oat grain yields but increased the test weight. Reductions in oat yields due to the presence of clover increased as oat row width increased. The yields were reduced 7, 10, 14, and 18 percent respectively for the 8, 16, 24, and 32 inch row spacing. Reduced yields of oats were attributable to the companion clover and were 17 percent in 1954, 7 percent in 1955, and 12 percent in 1956.

7.4 Effect of Harvesting Stages on Forage Yield and Quality:

In feeding ruminants the nutritive value of the roughage is important from the economical production viewpoint of meat and milk, the formulation of rations, the feeding and management of animals on pasture or on other forage systems for maximum meat and milk production, and accurate interpretation of performance records of animals fed rations containing large amounts of forage. The use of roughage as a feed for ruminants depends on the age of the animal and the purpose for which it is fed, the amount of roughage consumed, and its chemical composition, digestibility, balance of nutrients, and economy.

It has long been recognised that the stage of maturity at which a plant is harvested is one of the most important factors influencing its chemical composition and nutritive value.

Extensive reviews on the importance of roughage quality in ruminant rations have been made by Huffman (1939, 1953), Reid et al. (1959), Van Riper and Smith (1959), Sullivan and Garber (1947) and Stallcup et al. (1956) have reviewed the literature on the influences of the stage of maturity on the nutritive value of certain plants.

Newman (1894) studied the influence of stage of maturity of rye on the chemical composition of the plant under Arkansas conditions. In the period from the boot to the blossom stage, crude protein decreased and crude fibre increased. Rosen et al. (1953) studied the crude protein of oat forage from November through March. Early growth in the autumn was much higher in protein than were later clippings.

Thurman et al. (1957) have reported on the chemical composition of the oat plant from April through June. In general, crude fibre increased with advancing maturity and protein was reduced. Digestion trials were conducted on silages harvested at the boot, milk, and hard dough stages of maturity. The total digestible nutrients decreased from 71.3 to 58.8 percent from the boot to hard dough stages. Stallcup (1958) published a preliminary report on the change in composition of some forage plants at different stages of maturity.

The apparent influence of ash and the buffering capacity of plants on rumen pH has been reviewed by Cason et al. (1954). The influence of lignin on digestibility has been reviewed and data on some forages have been published by Stallcup et al. (1956). The composition of crude fibre in several forage plants has been shown to vary widely by Stallcup (1958). Meyer et al. (1957) made detailed studies of the changes in composition and nutritional value of oat forage with stage of maturity under California conditions. Austenson and Law (1958) undertook an extensive review of literature pertaining to the effect of fertilizer on chemical composition of pasture herbage.

Little information is available in Pakistan on forage yield and quality in oats, barley, and wheat crops harvested at various stages of maturity. Studies were thus carried out to determine an ideal maturity/harvesting stage to obtain a compromise between maximum forage yield and a reasonably good forage quality.

Hussain et al. (1998) evaluated oats, barley, and wheat for forage yield and quality at nine maturity stages at NARC, Islamabad during 1990 –1992. It was observed that oats harvested at head emergence stage, and barley and wheat at 100 percent flowering stage produced maximum green forage yields. In all three crops the highest dry matter yield was recorded at early dough stage. The maximum crude protein content was recorded at 4-leaf harvesting stage repeatedly, whereas the minimum protein contents were recorded in the early dough stage. Oats, barley, and wheat harvested at boot stage provided a good compromise of green fodder yield, dry matter yield, and forage quality. At this stage a sufficient quantity of fodder with moderate forage quality was obtained (Table 6). Recent work on oats is reported by Hussain et al. (2001).

Table 6. Green fodder yield, dry matter yield and crude protein contents of oats,

barley and wheat under various cutting stages

Cutting stages Green fodder yield (t/ha) Dry matter yield (t/ha) Crude Protein percent
Oats Barley Wheat Mean Oats Barley Wheat Mean Oats barley Wheat Mean
CS 1 37.66 33.15 21.73 30.84 5.81 4.80 3.93 4.85 14.93 13.47 12.56 13.65
CS 2 40.43 35.74 26.26 24.15 7.13 5.75 4.78 5.89 14.07 12.78 11.97 12.34
CS 3 56.45 44.77 24.49 41.90 10.68 7.49 4.67 7.61 12.65 11.70 11.53 11.36
CS 4 67.16 51.30 28.80 49.09 12.41 9.21 6.52 9.39 10.80 9.85 10.21 10.28
CS 5 69.44 48.45 31.78 49.89 12.15 8.05 7.41 9.21 8.75 8.42 8.15 8.44
CS 6 64.60 56.39 30.66 50.55 11.43 10.26 7.08 9.59 8.10 7.72 7.50 7.77
CS 7 68.21 58.42 40.51 55.71 13.35 11.17 9.83 11.44 7.63 7.54 7.32 7.50
CS 8 64.27 53.93 33.95 50.72 13.99 13.40 10.83 12.74 7.50 7.02 6.96 7.16
CS 9 51.21 42.52 33.72 42.52 17.17 14.67 12.23 14.69 7.15 6.85 6.75 6.92
Mean                 10.17 9.22 9.48  

Hussain et al 1998.

CS 1 = Repeated cuttings at 4-leaf, CS 2 = Repeated cuttings at tillering

CS 3 = Repeated cuttings at jointing, CS 4 = Repeated cuttings at boot

CS 5 = Harvesting once at head emergence, CS 6 = Harvesting once at 50 percent flowering

CS 7 = Harvesting once at 100 percent flowering, CS 8 = Harvesting once at early milk

CS 9 = Harvesting once at early dough

Hussain et al (1995) conducted trials during 1990 and 1991 to evaluate yield and quality of fodder at different harvesting stages on oats and barley (Table 7). It was concluded that oats harvested at booting stage and barley at 100 percent flowering stage produced maximum forage yield (79.45 and 63.10 tons/ha respectively). In oat and barley crops, the highest dry matter yield (15.54 and 13.75 tons/ha) respectively was recorded at early dough stage. In both the crops crude protein contents decreased with the advance in crop maturity. The maximum crude protein content (14.93 and 14.37 percent) in oats and barley respectively was observed when the crops were harvested repeatedly at the 4-leaf stages, whereas the minimum was at the early dough stage in both crops. Oats and barley harvested at booting stage proved better for reasonable forage yield (67.32 tons / ha), dry matter yields (11.66 tons/ha) and forage quality (crude protein 10.33 percent). At this stage a sufficient quantity of forage yield with moderate quality was obtained in both crops (Table 7).

 

Table 7. Green fodder and dry matter yields and crude protein of oat and barely as affected by cutting at different stages

Cutting stage Green-fodder yield (tons/ha) Dry-matter yield (tons/ha) Crude protein ( percent)*
Oats Barley Mean Oats Barley Mean Oats Barley Mean
CS 1 46.02 39.63 42.82 7.40 5.58 6.49 14.93 13.47 14.20
CS 2 51.39 42.04 46.71 9.23 6.91 8.07 14.07 12.78 13.43
CS 3 69.35 48.68 59.02 12.63 8.26 10.45 12.65 11.70 12.18
CS 4 79.45 55.19 67.32 13.96 9.37 11.66 10.80 9.85 10.33
CS 5 72.69 51.16 61.92 12.66 7.39 10.02 8.75 8.42 8.59
CS 6 70.98 54.95 62.96 11.48 10.00 10.74 8.10 7.72 7.91
CS 7 77.78 63.10 70.44 14.18 11.12 12.65 7.63 7.54 7.59
CS 8 73.26 58.22 65.74 14.09 11.70 12.90 7.50 7.02 7.26
CS 9 51.97 45.72 48.84 15.54 13.75 14.64 7.15 6.85 7.00
LSD(P=0.05) 10.47 7.40   2.94 2.08        

* Data for one year

Hussain et al 1995

CS1= Repeated cutting at 4-leaf; CS2= Tillering;

CS3= Jointing CS4= Booting stage;

CS5= Harvest once at ear emergence; CS6= 50 percent flowering;

CS7= 100 percent flowering; CS8= Early milk

CS9= Early dough stage.

Lawes and Jones (1970) investigated the ensiling characteristics of barley, wheat, and oats crops cut at two different dates and reported that dry matter yields increased until the end of July. Digestibility and crude protein content declined until mid July when values of 56-60 and 6-7 percent for the barley and wheat and 53-55 and 5-6 percent for the oats respectively were maintained. Crops of all varieties were suitable for ensiling at maximum yield.

Brundage and Sweetman (1967) have recommended that oat/pea forage be harvested in the late milk stages of oat maturity to take advantage of higher yields of lower moisture silage. Bolsen et al. (1976) and Polan et al. (1968) recommended harvesting small grain forages at the dough stages of maturity. Comparing oat forage ensiled at the boot, early milk, and soft dough stages of maturity, Martz et al. (1959) recommended harvesting at the boot stage or soon thereafter.

7.5 Oats as a dual-purpose crop:

Seed production is a major problem with the majority of forage crops as most of them are harvested and fed to livestock well before seed formation, unless intentionally kept for seed. Therefore, development and cultivation of dual purpose forage crops varieties could be one of the best options/solutions for seed production and availability. The most common cereal grown as a dual-purpose (harvesting/grazing plus grain) crop in Pakistan is oats.

Dost et al. (1994) evaluated oat cultivar S-81 under different maturity/ harvesting treatments during 1991-92 at NARC, Islamabad (Table 8). It was reported that forage yield, dry matter yield, and crude fibre increased while seed yield and crude protein declined by harvesting the crop with advances in maturity. Maximum forage and dry matter yields with considerably inferior quality forage (as determined by lower crude protein and higher crude fibre values) were observed at 50 percent flowering stage. Minimum forage and dry matter yields with superior quality forage were recorded in the crop harvested at 70 and 85 days after planting. The results of this research also indicated that it is possible to have a satisfactory forage yield, forage quality and grain yield from oats crop that has already been harvested once for forage at 115 days after planting.

Table 8. Average Plant Height (PH), Tillers/Plant (TL), Leaves/Tiller (LV), Green Fodder Yield (GY), Dry matter Yield (DY), Seed Yield (SY), Crude Protein Contents (CP), and Crude Fibre Contents of Oats cultivar S-81

Cutting treatments PH TL LV GY DY SY CP CF
CT 1 63.63 d 6.18 c 4.89 c 13.59 d 2.07 d 0.85 b 12.29. a 22.32 d
CT 2 83.70 c 6.80 ab 5.89 b 16.92 d 2.57 d 0.54 c 12.25 a 22.94 c
CT 3 90.06 bc 6.89 ab 6.26 ab 23.46 c 4.08 c 0.48 8.68 b 24.50 b
CT 4 94.74 b 7.07 a 6.59 a 33.33 b 6.60 b 0.46 8.46 25.58 a
CT 5 134.78 a 6.46 bc 6.56 a 54.99 a 12.99 a - 7.94 25.60 a
CT 6 - - - - - 1.34 a - -
LSD (0.01) 10.70 0.54 0.63 4.73 1.24 0.24 0.34 0.36
Means followed by the same letters do not differ significantly at 1 % probability level.Dost et al. (1994)

CT1= Cutting for fodder after 70 days of planting and then for seed,

CT2= Cutting for fodder after 85 days of planting and then for seed,

CT3= Cutting for fodder after 100 days of planting and then for seed,

CT4= Cutting for fodder after 115 days of harvesting and then for seed,

CT5= Cutting at 50 percent flowering for fodder only,

CT6= No cutting for fodder but for seed only.

Tilt (1965) reported that skilled grazing management of dual-purpose oats has a big effect on ultimate grain yield. Early sown oats can be grazed from the end of April through to August when they are either ploughed under or "shut up for grain". The later the grazing continues, the more the grain yield is likely to be depressed. In 1964, a plant breeding programme to develop a dual-purpose variety adapted to Tasmanian conditions was initiated by the Department of Agriculture and resulted in the release of "Esk" in 1975 for all Tasmanian areas (Vertigan 1975). "Esk" is now the most common oat variety in Tasmania, and is normally sown between February and April. It is used for winter grazing by sheep and, depending on the farmer’s requirement for stored (conserved) feed, may be harvested for hay or grain. It is late maturing, resistant to most diseases and to grain shedding (Vertigan, 1975).

Smith (1960) found that the analysis of oat forage indicated that the percentages of protein, fat, ash, P, Ca, K, and nitrate declined from early growth to maturity while the percentage of fibre increased until the heads were emerging from the boot, and then decreased. The yields per acre of dry matter and of protein, ash, P, and Ca, constituents important to animal nutrition, increased steadily to a stage of growth when the seeds were nearly ripe and then decreased. However, a high proportion of the total yield per acre of protein, ash, P, and Ca was produced at early dough, the stage of growth often indicated as the best stage to harvest for hay or silage.

Forage stand losses resulting principally from lodging of oats and volunteer growths from shattered seed have promoted widespread interest in the harvesting of oats at immature stages of growth as pasture, silage, or hay. Chemical analyses (Berry 1920, Fagan and Watkin 1931, Keitt and Tarbox. 1912, Meyer et al. 1957, Nicholson 1957, Smith and Ross 1943, Sotola 1937, Thurman et al. 1957, Underwood and Moir 1944) and limited feeding trials (Meyer et al. 1957, Sotola 1937) have shown that oats at immature stages of growth can provide nutritious and high quality forage. The most complete studies of the changes in composition of oats from early to late stages of growth have been made by Berry (1920) in Scotland and by Sotola (1937) in the state of Washington, USA. However, many of the analytical studies have been made at only the more advanced stages of growth (Fagan and Watkin 1931, Keitt and Tarbox 1912, Nicholson 1957, Thurman et al. 1957, Underwood and Moir 1944) or have been made for only a few chemical constituents (Henderson and Davis 1955, Meyer et al. 1957, Smith and Ross 1943).

Several investigations have shown that grazing or clipping of small grain stands can provide, in addition to grain, high quality forage and reduced lodging. However, clipping frequently decreased subsequent grain yields (Gardner and Wiggins, 1960, Crowder 1953, Cutler et al. 1949, Thakur and Shands 1954, Washko 1947, Welton and Morris 1931, Westrate and Grafius 1958). The effects obtained have varied with the soil and climatic conditions (Cutler et al. 1949, Sprague 1954), nitrogen fertilization (Morris and Gardner 1958), and with the date of clipping.

In Saudi Arabia, Farnworth and Williams (1977) planted oats, barley, and wheat crops for two cuts for forage followed by grain. The first cut for forage was obtained at a stubble length above ground of 2.5 cm and 12.5 cm. Oats produced the highest forage yields but lowest grain yields while wheat produced highest grain yields. Similarly oats produced higher yields of straw. It was also found that oats had higher protein, fibre, and ash levels than barley or wheat but lower carbohydrate levels. Barley had the lowest protein level and wheat the lowest fibre and ash levels.

7.6 Fertilization of winter forages :

Application of different doses of nitrogenous and phosphatic fertilizer produced substantially higher yields (at the five locations in Pakistan indicated in Table 9) than using no fertilizer or farm yard manure alone, especially on seriously depleted soils, which had been mono-cropped for many years with wheat/maize/ barley etc. Maximum forage yields were obtained through application of 150-75 N P kg/ha. at most of the locations under trial and was followed by 150-25 N P kg/ha (Table 9). However, increased usage of chemical fertilizer could not realistically be justified in many instances for economic and environmental reasons (Table 9).

Table 9. Green fodder yield of oats cultivar Scot (tons/ha) under different

fertilizer doses at various locations during 1999-2000

Fertilizer

N P kg / ha

Green Fodder Yield (tons/ha)
1

NARC Islamabad

2

ARI

Tandojam

3

ARI, Tarnab

4

ARI, Sariab

5

AARI Faisalabad

100-25 74.69 59.56 30.78 46.00 96.29
50-50 61.11 52.16 28.01 42.33 79.32
50-25 56.79 57.40 25.23 40.33 70.37
50-75 61.11 55.24 28.94 47.33 101.23
150-25 80.86 74.99 29.40 45.33 98.45
150-75 87.04 74.09 30.78 56.66 112.04
100-50 74.69 62.34 31.95 50.00 99.38
00-00 37.65 44.13 27.78 32.00 52.16
National Agricultural Research Centre, Islamabad

Agricultural Research Institute, Tandojam

Agricultural Research Institute, Tarnab, Peshawar

Agricultural Research Institute, Sariab, Quetta

Ayyub Agricultural Research Institute, Faisalabad

In India, Taneja et al. (1981) found that with the application of 120 kg N/ha to the oats and barley crops, the green fodder and dry matter yields increased significantly. For seed yields, however, the significant response was observed up to 80 kg N/ha. Also the application of 80 kg N/ha provided the maximum net profit of Rs. 4022.30/ha.

Available nitrogen is usually the limiting factor in dry matter production of winter forage (Crofts, 1966b), and oats and other small grain cereals have shown considerable increase in forage and grain yield due to nitrogen fertilization after grazing. The response will vary according to the temperature, soil type, soil fertility level, rotation system, irrigation, cultivar, date of sowing, time of nitrogen application and other environmental factors.

In USA, Ohm (1976) recommended that more extensive testing of potential new cultivars be undertaken to determine specific fertility x cultivar interactions. It was found that application of nitrogen resulted in significant increases in plant height, lodging score, percent protein, and yield in oat cultivars; it did not affect heading date or seed weight; and it reduced test weight. Short-strawed types increased in plant height more with nitrogen application than several taller cultivars although plant height increases were greatest for other tall types. Lodging scores of weaker-strawed cultivars did not always increase more with nitrogen application than those of stronger-strawed cultivars. Percent protein in certain low protein cultivars increased more than that of some high protein cultivars with fertilizer nitrogen. A given cultivar may not respond similarly to nitrogen in terms of percent protein and/or yield. More extensive testing of potential new cultivars to determine specific fertility x cultivar interactions was recommended.

In Australia, autumn sown oats has shown a large response in forage yield to nitrogen fertilization and irrigation, with economic benefits in reducing the winter-feed shortage (Crofts, 1966a, 1966b; Brown, 1975). Blackman (1936) showed that nitrogen can increase the growth rate of grasses when soil temperature at 10 cm is in the range of 5 -8 0C, as the slow release by ba