1. Introduction
2. Soils and Topography
3. Climate and Agro-ecological Zones
4. Livestock Production Systems
5. The Pasture Resource
6. Opportunities for Improvement of Fodder Resources
7. Research and Development Organizations and Personnel
8. References
9. Contacts

Niger (French République du Niger) is landlocked in the heart of West Africa. It is bordered by Algeria and Libya on the north, Mali on the west, Burkina Faso and Benin on the southwest, Nigeria on the south and Chad on the east (see Figure 1). It has a total area of 1,267,000 sq km, but only half of this is habitable due to adverse climatic or soil conditions. Niger is divided into one capital district and seven departments, which are subdivided into districts and communes. Niamey is the capital, and largest city; Zinder, Maradi, Tahoua and Agadez are the other principal towns. French is the official language, but Haussa is the language of local trade. Other African languages, such as Fulfulde, Tamachek, and Djerma are also used extensively.

Figure 1. Map of Niger

Since the end of the Stone Age different groups began to settle in the more humid environments around the Air massif, along the borders of the former mega-lake Chad, and in the Niger valley. The knowledge of writing as one factor of cultural development did not appear before the arrival of Islam in the eighth century with the Arabs. Royal empires of unknown age already existed at that time. Finally, after their power decreased in the 19^th century Niger was a French military territory and then became a colony in 1922 under a military regime. On 3rd August 1960, after elections, Diori Hamani became the first president of the independent Republic of Niger.

Only 29 percent of primary school-age children receive education. In 2001 the average life expectancy at birth was 42 years; the infant mortality rate was 124 deaths per 1,000 live births (2001 estimate). According to the World Factbook the population in July 2006 was estimated at 12,525,094 with a 2.92% annual growth rate. More than 60 percent of the population lives below the poverty line.

Economy. A particularly harsh climate, inhospitable geographical features and a completely land-locked situation, such are the underlying economic realities of the Republic of Niger. However, subsistence farming and stock-rearing contribute approximately 40 percent to the GDP (second only to services, which provide 42 percent) and nearly three-quarters of the labour force is employed in this sector. Based principally on agriculture and livestock breeding, the country's economy is still largely at the mercy of the vagaries of the climate. Niger has tin, gold, and uranium mining operations. Large deposits of uranium are located in northern Niger and still, it counts for about three quarters of export earnings. Salt is also mined in Niger in small quantities. Industry is limited mainly to food processing and construction, manufacturing enterprises are mostly very small. Burning wood and other traditional fuels accounts for 80 percent of the country's energy consumption.

Agriculture. Less than 4 percent of the country is arable, 9 percent are permanent pastures and only 2 percent are forests and woodlands. Agriculture is limited in the north by the 350 mm isohyet beyond which the millet (Pennisetum glaucum) production practically ceases. The Nigerien Sahelo-Sudanien and Sudanian zones are considered as potentially sedentary areas (in contrast to the northern zones where primarily nomad activities are found). Here, mainly millet (Pennisetum glaucum) and Sorghum (Sorghum bicolor) - traditional extensive crops and covering most of the cultivated soil - cowpea (Vigna unguiculata) and groundnut (Arachis hypogaea) are found. Some cotton (Gossypium spp.) appears further south, in the Sudanian zone. The depressions of recent and ancient rivers are for vegetable (tomatoes, onions, etc) growing or some fruit trees. Rice is cultivated around the river Niger and river Komadougou. Other crops of lesser importance are sugar cane, maize, and sweet potato.

The yearly production is presently approximately 3,500,000 tons of cereals, supplemented by thousands of tons of international aid. The mean yield is around 400 kg/ha for millet and around 190 kg/ha for sorghum, but there are big differences between the regions. Cowpeas, cotton and groundnuts are cultivated for export. Millet (Niger is the world's top producer per capita), sorghum, cassava, pulses, rice, sugar cane and some vegetables are grown for local consumption. Fishing is conducted in Lake Chad and the Niger river, and the catch is consumed or exported locally.

The share of cultivated land is increasing and competes severely with livestock keeping. In fact, livestock keeping is now limited to the northern part of Niger: domestic animals are widespread, staying the whole year or moving one part of the year to northern zones. Agriculture benefits from organic fertilization that maintains soil fertility in the vicinity of settlements. In search of an increase of arable soil, woodland is cleared. Clearing and wood-exploitation reduces considerably the original vegetation: just for the capital city Niamey, more than 11,000 tons of firewood per year are needed. Additionally, fallow land is lessened and rotation brought down to 2-3 years, further diminishing soil fertility.

Wildlife. About 7.7 percent of the Niger's land is officially protected. Although Niger has banned hunting (except for scientific purposes) throughout the country, rampant poaching seriously threatens wildlife populations. But other reasons also play an important role in the decline of the Nigerien wildlife population: the destruction of the habitat by agriculture, deforestation, fire, and the competition with domestic animals. Natural events such as the heavy droughts have also reduced the pasture grounds for wild animals.

In Niger, the principal fish habitats are the Niger river and lake Chad. Some fish species are adapted to an environment with no water for a certain time by surviving dryness in a torpid state inside the sediments or covered by a mucous protection membrane or in the egg stage. A few fishes have locally a considerable economic importance (e.g. Nile perch, Lates niloticus). Amphibians in Niger are generally bound to permanent water places, but some species have a more terrestrial life and survive the long dry season by a very short life cycle during the rainy season and/or by digging themselves into wetter soil layers.

Among the noteworthy Nigerien reptiles are species protected by international conventions such as the Nile crocodile (Crocodylus niloticus) and lizards (e.g. Varanus griseus, Varanus niloticus). Interesting species are also Python snakes (Python sebae, P.regius), water turtles (e.g. Trionyx triunguis, Pelomedusa subrufa) and tortoises (e.g. Testudo sulcata). More than 600 bird species are found in Niger, many of which are threatened. Among the most interesting birds are eagles and vultures (Accipiteridae), ground-hornbills (Bucorvus abyssinicus), and the ostrich (Struthio camelus). Guinea fowl (Numida meleagris) and pigeons (Columbidae) are frequent and embellish the daily rural diet. Mammals are listed with 131 species and numerous species are endangered. None of the mammals are endemic. Important species include antelopes such as the Scimitar Oryx (Oryx dammah), the Addax (Addax nasomaculatus), the Dama gazelle (Gazella dama) or the Slender-horned Gazelle (Gazella leptoceros). Conspicuous but rare mammals are the African Elephant (Loxodonta africana), the Cheetah (Acinonyx jubatus) and the Lion (Panthera leo). Hippos (Hippopotamus amphibius) inhabit the waters of the Niger river. East of the Niger river are areas with a few nomadic giraffes. In Nigerien savannas relatively frequently encountered are warthogs (Phacochoerus africanus), monkeys such as Baboons (e.g. Papio cynocephalus). Hyenas (Hyaena hyaena) can be found all over Niger.

The Nigerien government tries to preserve animal lives in reserves such as the 'W' National Wildlife Park (220,000 ha - also an important tourist attraction) and the Tamou Nature Reserve (78,000 ha) which are located 125 km south of the capital Niamey. In the southern northern/northern Sahelian zone, the Aïr and Ténéré Strict Nature Reserve (7,700,000 ha), and within it, the core Addax Sanctuary Strict Nature Reserve, were set up in 1988. The latter closed area protects the very rare Addax from the vehicles of hunters and tourists who formerly chased them.

Niger is a vast, undulating plain at an average altitude of 300 m above sea level. There are, however, a number of extensive depressions, usually filled with sand, the tops of the ridges sometimes being formed from volcanic structures, the highest of which is Mount Greboun (2,310 m) in the Aïr Massif. The country can be roughly divided into three zones: the north, centre and south. The northern zone, covering about two thirds of the surface area, is in the Sahara. It is an elevated region formed by plateaux and mountains, and with the exception of some isolated oases, vegetation is rare. The centre is part of the Sahel (the Ténéré region). It is a semi-arid region with few trees. The south is the only fertile and wooded region and where it rains enough for food crop farming without irrigation.

Apart from a few more or less permanent pools and seasonal water courses, Niger's main hydrographic network consists of one major river, the Niger, a smaller river, the Komadougou Yobé, and Lake Chad. The river Niger crosses the country in the south-west over a distance of 550 km. The river Komadougou Yobé marks the boundary between Niger and Nigeria over a distance of 140 km. The Nigerien part of Lake Chad covers some 3,000 square km. The lake itself is the vestige of an ancient quaternary sea, at an altitude of 280 m and today at no point is more than 4 m deep. Islands proliferate in the lake and the evaporation rate is extremely high; 98 percent of its water supply comes from the Chari River and from rainfall.

In the extremely dry areas in the north and northeast of Niger, scarcity of water and vegetation inhibits soil formation. Some parts have, however, been subject to mechanical fragmentation. Strong winds take away the finer parts, sand and silt. The A horizon is little or not developed and the organic matter content is generally less than 0.5 percent in the upper 40 cm (in the former World Reference Base (WRB) of FAO, many of these soils have been classified as Yermosols, but this soil group was deleted in the revised WRB. Since, the soil map of Africa has not been updated, the name Yermosols will therefore be retained here. According to the revised WRB, these soils would now, depending on certain properties, belong to Calcisols, Cambisols, Durisols, Gypsisols or Arenosols). In mountainous regions (Aïr or the Djado plateaus), rocks are covered by a shallow and coarsely fragmented soil layer (Leptosols). Productivity is low and these soils can be used only for extensive grazing (see Figure 2).

Figure 2. Soil map of Niger [Click to view full map]

Further to the south a large strip of sandy soils crosses Niger from Mali to Chad. The slightly weathered soils, Cambic Arenosols in the eastern part, show even thin layers of clay accumulation in the central and western part (Luvic Arenosols). As rainfall and thus weathering becomes more regular, soils start to show a light coloured upper horizon with little organic matter, and a reddish lower horizon due to a higher iron-concentration. Arenosols have a low water retention but are permeable and easy to work. They have a low natural fertility, a low cation exchange capacity and deficiencies of minor elements are common. These soils may erode easily by wind and water as soon as vegetation is gone. They are suitable for little-demanding agricultural crops such as millet or groundnut.

On the terraces between the Niger river and the Dallol ('valley') Bosso and the Dallol Maouri, in the south-western part of Niger, the Arenosols are associated with Regosols, i.e. soils on loose parent material that are almost without soil development. The hydromorphic soils, with a high groundwater, in the depressions of south-central Niger (Tarka valley, Goulbin Kaba and Goulbin Maradi) and at Lake Chad, and the soils on alluvial deposits (Fluvisols), generally offer good and intensively used agricultural soils. Cambisols (moderately weathered soils) and Vertisols (heavy clay soils) are only of limited extent in Niger.

Niger is situated in one of the sunniest regions of the world, and has a mainly dry climate with considerable temperature variations. Potential evaporation is 2 to 4 m per annum, while rainfall nowhere exceeds 800 mm and even falls to below 100 mm over almost half the country. The rainfall pattern is Saharan in the north, with some 160 mm falling in less than one month, except in the desert where it practically never rains, and Sudano Sahelian in the south, where some 600 mm of rain falls during three to four months (June-September). Rainfall varies, however, from one region to another and its distribution is very erratic with levels falling sharply as one moves northwards. Temperatures can exceed 40 degrees Celsius in the shade from March to June, period of the 'harmattan' wind. During this period come the first rains usually announced by heavy clouds moving across the sky. From November to February, temperatures drop considerably, particularly with nightfall. Annual temperature differences vary from around 16 ° C in the northeast to around 9 ° C in the southwest.

Vegetation. Niger covers two geobotanical zones: the Saharo-Sindian in the very north and the major part, the Sudano-Zambezian zone. Biogeographically Niger covers three areas: the Saharan, the Sahelian and the Sudanese. The transition between them is not clear-cut. Within tropical Africa, Niger presents one remarkable characteristic: the Ténéré (in the northern part of Niger) - one of the world's floristically poorest zones: there are fewer than 10 different species per 10,000 square km.

The Saharo-Sindian zone in the north: little and very irregular, or sometimes no rain. The dominant vegetation of the plains, if there is one, is a discontinuous grassland (often referred to as "steppe" in French publications) which is mostly limited to depressions. It belongs to the Acacia-Panicum grouping, dominated by Panicum turgidum. Plants are adapted to the absence of water. On the dunes Cyperus conglomeratus is abundant in the herbaceous carpet associated with various Poaceae, like Aristida mutabilis, Tragus racemosus, Cenchrus biflorus or Lasiurus scindicus. On the plateaux Aristida hordacea, Aristida funiculata, Cenchrus ciliaris, Eragrostis pilosa and Schoenefeldia gracilis are common. In the dry valleys, Cymbopogon schoenanthus is dominant in the grass carpet. The very scarce woody layer includes, especially, Acacia raddiana, but also Acacia ehrenbergiana, Cordia sinensis, Maerua crassifolia and Grewia tenax. Variations in the vegetation are mostly due to small changes in the micro-climate, the soil or relief. Thus, the formations of the mountains of Aïr show substantial differences compared to those of the plains. The formations of the lower stage of the Aïr (between 500 and 900 m of altitude) have woody stands made up often of Maerua crassifolia, Leptadenia pyrotechnica, Calatropis procera and Salvadora persica that are associated with Balanites aegyptiaca and various Acacia. Among the grasses, Stipagrostis spp., Aristida spp. and Eragrostis spp. are common. On the higher levels of the Aïr ( > 900 m) formations of Acacias (Acacia raddiana, Acacia ehrenbergiana, Acacia laeta, Faidherbia albida 'Gao') with Boscia senegalensis, Commiphora africana, Balanites aegyptiaca, Ficus cordata, Hyphaene thebaica, Ziziphus mauritania, Anogeissus leiocarpa, Bauhinia rufescens and Salvadora persica are found. The Poaceae are represented by Aristida mutabilis, Cymbopogon schoenanthus, Tripogon multiflorus and Desmostachya bipinnata.

The Sudano-Zambezian zone is divided into two parts: the Sahelian belt in the north and the Sudanian belt in the south. The Sahelian belt shows steppe vegetation whose woody layer thins out from a cover ranging between 15 and 60 percent in the south, to a shrubby aspect in the north. The grass layer is made up mainly by annuals. The Sahelian belt is again subdivided into two economic sectors, roughly at the 350-mm isohyet: A northern sector (Sahelo-Saharan), known as a zone of transhumance, because the cultivation of millet is hardly possible, and a southern sector (Sahelo-Sudanien), the sedentary zone, with many villages (where, however, cattle are still very numerous but away during the rains on a transhumance towards the north in order not to damage the crops). In all subzones, vegetation is distributed following the pattern of three major soil types: sandy soils, soils with sufficient clay and silt content and soils on outcrop parent material.

The woody species of the sahelo-saharian sector are mainly of trees such as Faidherbia albida, Sclerocarya birrea, Boscia senegalensis, Balanites aegyptiaca, Hyphaene thebaica, Acacia raddiana and shrubs like Salvadora persica. The grass layer is made up of Cenchrus biflorus, Cenchrus prieurii, Brachiaria xantholeuca, Brachiaria villosa, Schoenefeldia gracilis, Panicum turgidum, Aristida mutabilis and Eragrostis tremula. In the eastern part of this sector, i.e. on the sandy alluvium of the Tal desert or the clayey plateaus of the Kadzel, the woody layer includes Acacia raddiana, Piliostigma reticulatum, Balanites aegyptiaca, Commiphora africana, Grewia villosa, Grewia tenax, Guiera senegalensis, Combretum micranthum, Acacia senegal, Acacia nilotica, Hyphaene thebaica, Sclerocarya birrea, Piliostigma reticulatum and shrubs like Leptadenia pyrotechnica and Calotropis procera. The grass layer is mainly composed of Cenchrus biflorus ('cram-cram'), Cenchrus prieurii, Tragus racemosus, Eragrostis tremula, Dactyloctenium aegyptium, Aristida funiculata and Aristida adscensionis.

In the Sudano Sahelian sector, trees are commoner and even thickets are found. The steppe vegetation persists on sandy soils of dry valleys with trees such as Guiera senegalensis, Acacia nilotica, Combretum micranthum, Piliostigma reticulatum, Prosopis africana, Faidherbia albida, Acacia senegal, Ziziphus mauritania, Acacia raddiana, Balanites aegyptiaca, Hyphaene thebaica and Boscia senegalensis. The grass layers of these steppes are rich in Aristida adscensionis, Eragrostis tremula, Bracharia villosa, Cenchrus prieurii, Cenchrus biflorus, Aristida mutabilis, Aristida funiculata, Enteropogon prieurii, Dactyloctenium aegyptium and Schoenefeldia gracilis. Plateaux carry savanna formations and thickets occupy the southernmost Sahelian belt and extend on the lateritic surfaces covered with thin soil . Shrubs and trees of Combretaceae, Tiliaceae and Mimosaceae form what is called the 'tiger bush' and 'spotted bush' in which stripes of woody vegetation and bare ground alternate. The grass layer contains, for example, Aristida adscensionis, Elionurus elegans, Eragrostis tremula, Pennisetum pedicellatum, Schoenefeldia gracilis, Tripogon minimus, Brachiaria xantholeuca, Dactyloctenium aegyptium, Eragrostis tremula and Eragrostis pilosa. These forest ecosystems are a major element for livestock holding in Niger.

The Sudanian belt is relatively small in Niger, mainly in the west of the country and south of Niamey. It is more wooded than the Sahel and carries a vegetation characterized by an uninterrupted herbaceous layer where perennial species of larger size dominate. The grass layer is heterogeneous and particularly developed in clearings. Common species on lateritic plateaux are Loudetia togoensis, Brachiaria xantholeuca or Digitaria gayana; in the valleys or on gentle slopes, Andropogon gayanus and Pennisetum pedicellatum are frequent. The woody vegetation is characterized by a great number of sometimes dense stands of Combretaceae, and by the presence of species useful to man, like Butyrospermum parkii (the 'shea tree') or Parkia biglobosa ('néré'). The woody layer of variable density can form large stands, like open forests with Anogeissus leiocarpus, Bombax costatum, Acacia macrostachya, Daniella oliveri, Piliostigma reticulatum and more. Gallery forests that border the major rivers have a tree layer which can reach 20-25 m height and include trees like Mitragyna inermis, Cola laurifolia, Afzelia africana, etc.

North of the 12^th parallel of latitude, most of the south-west of Niger is covered by savannas with a woody layer dominated by Combretaceae and trees such as Faidherbia albida, Bombax costatum and Sclerocarya birrea. The grass layer is very often formed by Andropogon gayanus, Aristida spp. , Loudetia hordeiformis, Panicum nigerense and local spots of Cenchrus biflorus. In some valleys (i.e. Dallol Maouri), depending on the ground water, trees like Prosopis africana or Khaya senegalensis are found and palms such as Hyphaene thebaica or Borassus aethiopum can be frequently seen.

Livestock production contributes to 35 percent of the agricultural gross domestic product (GDP) and thus 12 percent of the total GDP (1995). More than one million people are working full time in livestock production, and more are in the agro-pastoral sector. After uranium, it is the second largest export product of Niger, but its share dropped from over 20 percent to less then 7 percent in 1995. Nigerien livestock production is based on extensive grazing but climatic vagaries, the extent and quality of pastures, sanitary and economic constraints set limits to its performance. The precarious situation favours two kinds of livestock:

- Camels excel because of their excellent adaptation to a dry environment; only they can exploit the entire Sahelian zone, especially the northern parts. Economically, it is a long term investment. By this strategy, the number of camels have increased by 50 percent since the severe droughts of the seventies and eighties whereas the number of cattle dropped by 20 percent in the same period .

- Small ruminants are a short-term investment and their numbers have doubled since the 1960s, increasing from 18 percent to 33 percent in numbers of TLU [one TLU = Tropical Livestock Unit = 250 kg of liveweight = 1 dromedary = 1.25 bovine or horse = 2 donkeys = 10 small ruminants]. In particular the number of sheep (see Table1) have increased (more than 70 percent in the last twenty years), although the number of goats is currently still superior (sheep 13 percent and goats 20 percent of the Nigerien total TLU). Due to their prolificity , small ruminants are favoured since a drop in number after a drought can quickly be rebuilt. Besides, they have a higher productivity compared to big ruminants.

In recent years, livestock production has shifted more and more into the agropastoral zone (and the agropastoral zone has extended into the pastoral zone), except for the extensive livestock keeping of the Bororo zebu. It is estimated, that approximately 2/3 of the bovine livestock are in the agropastoral zone today. Especially in the south-west of Niger, livestock fattening and dairy production is more and more intensified.

The use of complementary feed or additives depends on the level of livestock production. In the traditional domestic production in the villages, harvest residues and pod shells from field (cowpea) or tree legumes (Acacia, Tamarind, etc.) are used. In improved livestock production systems, close to urban centres and encouraged by international programmes, bran, brewer's residues or oilseed cakes are sometimes used. The volume of veterinary medicaments is difficult to estimate since there is an extensive black market but it is estimated to be below 0.05 percent of the economic value of the entire livestock.

Generally, milk is much liked in Niger and it constitutes the staple diet of people like the Peulh, Tuareg, Toubou and Arabs. Although Nigerien breeds of cattle such as the Azawak or the Kouri rank among the best milk cows in West Africa, the actual production is limited by malnutrition. For the most part, domestic milk production is for the private use of the livestock owner and less than one percent of domestic milk production is commercialised. Domestic milk production does not meet the demand, and milk imports amounted to more than US$ 13,250,000 in 1995 (38,000 tonnes of milk equivalent). On the other hand, the amount of milk available per capita dropped from 168 litres in 1968 to 37 litres in 1994. Industrial dairy production is predominantly based on imported milk powder.

Although the numbers are uncertain, it is estimated that in 1995 approximately 200,000 bovines, 30,000 camels, 600,000 sheep and 1,300,000 goats were slaughtered for consumption. Another important sector of Nigerien livestock marketing is hides, skins and leather, especially from goats. Estimates are 188,000 bovine hides, 85,000 sheep skins and more than 1,600,000 goat skins in 1994. Numbers of livestock by Department (1991-2000) are given in Table 1 and livestock numbers, meat and milk production, live animal exports and milk imports for the period 1996-2005 (FAO database) are given in Table 2.

Figure 3. Development of population and livestock in Niger  since 1991 [Click to view full image]

Table 1. Number of Animals per Department (1991-2000)
[ click here to view table ]

Table 2. Niger statistics for livestock numbers, meat and milk production,
live animal exports and milk imports for the period 1996-2005

Source: FAOSTAT 2006; n.r.  no record

* Other meat production in 2004: chicken meat 28,960 tonnes; meat of asses 1,760 tonnes; camel meat 7,800 tonnes and game meat 15,000 tonnes. 

** In 2004 goat milk production was 105,000 tonnes.

Figure 4. Niger - agricultural zones and important livestock markets [Click to view full image]

Livestock trade is organised through strong traditional networks linking producers, intermediaries, and traders. Most animal sales by producers take place in local markets. Nigerien urban centres and towns attract livestock for local consumption, but most is destined for export, mainly to Nigeria. The principal entry points to Nigeria follow major roads and rivers, even though animals are typically trekked across the border to avoid detection by customs agents. Table 2 provides details of live cattle, sheep, goat and ass exports from 1996-2005

Ethnical Background and Transhumance. In Niger, the stockbreeders belong primarily to two ethnic groups, the Peulh (Fulani) and the Touareg. The Peulh are present throughout the Sahel and nearly all over Niger (8 percent of the population) but nowhere are they a majority. The principal areas of their presence in Niger are, however, the south-western part, an area circumscribed by Dakoro, Agadez, Arlit and Tchin-Tabaraden, in an area between Zinder and Gouré; and finally in the south-eastern part, in the Manga region north of Diffa. The migrations of nomad Peulh in the northern pastoral Sahelian zone - it might be more accurate to call them infiltrations - are relatively recent.

It is necessary to distinguish the Peulh with their agropastoral activities from the exclusive Peulh stockbreeders who generally live further to the north. The nomad stockbreeders are most generally called Bororo, but name themselves Wodaabes. During the dry season, the Wodaabes are dispersed at many water points, quite far from each other. With the first rains, the camps move towards the first water holes, before gathering around the ponds and the new grass for the great annual regrouping ('worso'). The extent of this movement is not very significant and seldom exceeds 150 km. It is thus transhumance on a rather small scale which exploits water and fodder resources that allow more or less strong concentrations of cattle: at some ponds during the rains until sometimes December, to some wells supplied by a shallow water table between December to February. Then from March until the return of the rain, they move to places equipped with deeper wells with pumping devices. During the tough months (February to the beginning of the rains), herd-splitting becomes an important survival strategy.

Peulh are predominantly cattle breeders but it is the small ruminants that provide red meat for a family while cattle are capital investment and prestige. Within the family, the herding responsibilities are shared among different age and gender groups. Usually, the management of the herd is done by the men and children, while the culinary responsibility and food processing falls on the women, but they all learn the herding skills, too. Most Peulh continue herding well past middle age. The family owns the herd, although every animal has a designated owner. Meat surplus to immediate requirements may be dried, for later use, of for sale. The meat is cut thinly, beaten even thinner and spread over bushes. The Peulh show a particular behaviour ('pulaaku') that links them beyond economic differences. For example, the institution of solidarity of 'the attached cow' (habbanae), makes it possible to reconstitute a lost herd. The lender allows the borrower to take one of his heifers, three years in general, until this one has calved between one to three times, according to previously drawn up clauses. These calves belong to the borrower who will bring back the cow after the weaning of the last calf.

The agropastoralist Peulh occupy the southern Sahelian space. But even the transhumant Peulh groups are not strictly limited to stock-rearing: they usually sow millet near the fringe of cultivation during their migration. It is difficult to define exactly the share of agriculture and stockbreeding, and the activity of each social group is variable. In some cases old men and adults cultivate, whereas the young people lead the herds. In other cases entire families move with the herds while the head of the tribe stays with the fields. In all cases, stockbreeders have to move their herds far away from the cultivated zones during the rains, except for some calves and a dairy herd that remain in the villages. While it is essential for the agropastoralists to have the animals moved from the zone of cultivation, it is important for them to leave the areas of ponds infested by mosquitoes and midges, and to let the dry season grazing places recover.

The Touareg (or rather Kel Tamajeq as they call themselves according to a linguistic criterion) have a very feudal society, with nobles, clergy, vassals, artisans, and one-time slaves. They make up a group of equal importance (8 percent) to the Peulh, but unlike the Peulh, they are a majority in all the lightly populated pastoral areas west of a line joining the Eastern edge of the Aïr massif to Gouré (east of Zinder).

The Touareg habitually live in relatively small clans, that for ad-hoc defence of some common interest, can be merged quite easily with each other. Agriculture was initially practiced only in time of need and considered as a degrading activity. Now, Touareg are divided into many groups: some are still in transhumance and are exclusively stock breeders; others are still part of a pastoral economy, but stay in villages or camps close to their fields or their gardens; others add caravanning to their activities.

The exclusive stockbreeders occupy, in general, the land that is unsuitable for rainfed crops, to the north of the agropastoralists. Those among the Touareg who still live in the Aïr massif (the 'Kel Owey') are an exception since they can practise irrigated agriculture thanks to a high water table in the principal valleys of the Aïr.

Further east, the Touareg (called ' Iwellemmeden Kel Denneq') occupy a territory that goes from the agricultural zone to the desert according to a transect south-north without the obstacle of a border. During the dry season, these families move to different places in the south but during the rains, most of them converge on the area of Tegidda with its springs. Thus, the extent of transhumance is variable but generally longer than for the Peulh, sometimes close to a total of 1,000 km. More and more frequently, however, families stay at their fields and only some herdsmen and the young move with the herd.

The Touareg, who live in the sedentary zone of Birni' N-Koni, Madoua and Dakoro (the 'Kel Geress') sometimes have large herds of sheep and camels, which are taken in the dry season to the fallows of Nigeria (area of Sokoto) and in the rains to the springs of Tegidda. Those living in the north of the area (see section 5 below) of the 'salt cure', (in Abangarit, Tamesna), often have big herds which, in the dry season, live on pastures with 'alwat' (Schouwia thebaica) which exempts them from being watered. During the rains , they move in the opposite direction, to the south, to the area of Tegidda. The Touareg in the western departments (Niamey and Dosso) lost their mobility and are in an agropastoral economy. The transhumance of their herds is more and more opposed and limited by the extension of cultivated land and the presence of national borders.

The Toubou are only a marginal group of stockbreeders who live in eastern and north-eastern Niger. They are of a low numerical importance since they do not reach one percent of the native population of Niger. The Toubou call themselves Teda, when mainly involved in camel-raising or Daza, if cattle are the primary livestock.

Among the other stockbreeders, it is necessary to mention the Arabs. They form a small community in the east of Niger and north-west of Tchin-Tabaraden, but play an important role in camel-raising.

Livestock
In that part of the country north of the 350-mm isohyet which is unsuitable for agriculture, livestock are usually the primary source of food and income. However, livestock make an important contribution to the livelihood of the agropastoral farmers south of the 350-mm isohyet and are a source of insurance against some shocks brought by weather and economic disruptions.

Cattle breeds in Niger:

1. The long-horn zebu is mostly of the Bororo breed (approximately 60 percent of all bovines) and the Peulh type (10 percent). The Bororo breed (zebu bororodji) is a relatively tall animal with bulls of around 1.50 m in height and a weight of 300 - 400 kg. The female is smaller. The horns are lyre-shaped and the colour of the animal black or flame red. The skin has short shiny hair. The big ventral part of the belly is characteristic. The development of zebus is rather slow and it takes up to six years to complete growth entirely. At birth, calves weigh around 15-20 kg and the annual growth is around 70 kg per year. Lactation is usually six months and milk production relatively low with 3 - 4 litres at the beginning and 1.5 litres at the end of lactation. It is a fierce animal but it is very robust and suitable for transhumance since its body allows it to cover long distances. The animals are only slightly fattened and the meat yield is not more than around 45 percent.
2. The initial area of distribution of the Peulh breed (zebu djeli) was the Niger river, but it can now be found all along the southern border up to Lake Chad. It is usually white with crescent shape horns, smaller than the Bororo type and with a pronounced hump and ventral part of the belly. It is mostly reared by Peulh who practice a very limited transhumance and it is a better meat producer than the former type. Usually milk production is fair with around 2 litres per day for six months.
3. The short-horn zebu makes up approximately 25 percent of the cattle, and consists mostly of the Azawak type which is found on the whole Nigerien territory. The height of the Azawak bull is around 1.20 to 1.30 m, the weight around 300 - 350 kg, for the cow 200 - 250 kg. It is often of wheat colour but white, black, black and white, and other colours are also found. The horns of the bulls are short and thick; the cow has crescent shaped and thinner horns. It is a good animal for meat production that fattens easily, but the yield is quite low. It is also a good milker, under favourable conditions it produces for 7 - 8 months a year between 6 - 8 litres per day at the beginning and 2 litres at the end of lactation. The Azawak is normally a calm animal that can also be used for animal traction. It is especially found in the Touareg herds, and in the herds of the sedentary Hausa and Bouzou.
4. Finally, as a matter of interest, the Kouri taurin (Bos taurus typicus) should be mentioned. This animal, actually a good milk and meat producer, can still be found in small numbers around the Lake Chad. It has however some importance since it is an ancestor to many hybrid varieties.

Three breeds of sheep (with many crosses) are the most common in Niger.

1. The Peulh sheep has two principal breeds. The Bali-Bali is usually white with black or yellow spots around the eyes or ears. It is found in the northern zone since it does not tolerate the humidity of the Sudanian zone. The Oudah type also has a quite characteristic skin: A very clear-cut black or brown front and a white behind. A few animals are entirely white and obtain a high price at the time of Islamic festivities. This breed is typically part of the Peulh herds north of the zone of Trypanosomiasis. The size of the male Peulh sheep is around 80 - 90 cm in height and it weighs between 40 - 45 kg, whereas the female is smaller and weighs around 35 - 40 kg. The ram has strong spiral-shaped horns, while the simple horizontal horns of the female are sometimes absent. It fattens easily. At birth, the lamb weighs some 3 kg but reaches more than 8 kg after the first month and over 25 kg after 6 months. The milk production of the ewe is poor, around 0.2 litres per day during 6 months.
2. The Ara-Ara breed is generally found in the western part of Niger, from the south of the Saharan belt to Niamey. The male animal is between 80 - 90 cm in height and weighs around 40 kg; the female is between 60 - 70 cm and weighs around 30 kg. The colour is tawny or white sometime with black spots. The ears are short and the strong horns are spiral-shaped and much smaller for the female. The neck of the ram has frequently a roll of flesh. It is mainly the sheep of the Touareg pastoralists with herds of small ruminants. It produces a high quality meat and the milk production under good conditions can reach up to 0.6 litres per day during 6 months. The milk is consumed raw or transformed into a nationally famous cheese, the 'Tchoukou'.
3. The presence of the Koundoum race is limited to the vicinity of the Niger river and the Kolo islands close to the Malian border. It is much smaller (around 60 cm in height and 30 kg for the male), with a bare head, members and belly. The fleece is black, or white with black or tawny spots. This race was formerly used for wool production by the sedentary Djerma of the area. Although this race is usually not reared any more, it is often crossed with other breeds. The meat production is low but of high quality.

Goats are an important source of meat for nomads. Equally important is the production of hides. Two races are distinguished:

a) The Sahelian breed (with the Bouzou and the Peulh varieties) is found in most parts of the country. The buck is around 80 cm in height and weighs around 25 to 35 kg (the goat 75 cm and 30 kg). It is tri-coloured: black, white and red. The hair is very short, except for some withers with the buck. The meat production is low. The lactation is about 6 months and the goat produces an average of 0.6 litres per day.

2. The Red Goat of Maradi (known as Sokoto Red in Nigeria) is - as the name indicates - widespread in the area of Maradi. It is exclusively part of a sedentary livestock system, since it is not adapted to the hardship of the Sahelian climate and transhumance. It is a rather small animal (60 cm in height with a weight of 25 kg) with a beautiful chestnut colour and short shiny hair. Horns are only small. The milk production is considerable: the goat produces for more than 6 months around 0.7 litres per day and is, for some villages without bovines, the only milk source. Meat yield is good especially among the castrated males. Its real importance, however, lies in its worldwide highly appreciated leather quality.

Camel breeding plays an essential part in the life of the people in the north of Niger. Several breeds are known, i.e. the 'River' camel, the camel of 'Tibesti', the camel of 'Manga', the camel of Aïr or the 'Toubou' camel . Camels are used not only as a means of transport, but also as a source of meat and leather, and are often exported to neighbouring countries (Libya, Algeria).

The number of horses in Niger is considerable compared to other West African countries. Most breeds are derived from Barbs, Arab or Dongolow horses. The common breeds are the Djerma, the Hausa, the Songhoïand the Bozzan horse from the area around the Aïr. Horses are almost exclusively used for transport. Other animals of transport in Niger are donkeys , which in some areas are also used for animal traction in agriculture.

Swine raising is, for religious reasons, very limited and mostly for foreigners.

Poultry production plays a prominent role in the everyday meat supply. Chickens and Guinea fowl are the most important species. West African fowl breeds are innumerable, but generally, they are small and of little weight. Over the last decades, many American and European breeds have been introduced and crossed with local ones, i.e. Sussex , New Hampshire or Rhode Island Red

Animal health. Nigerien livestock and poultry are hit by many diseases among which the most important are:

- parasites: internal parasites (roundworms, tapeworms, hookworms, coccidia, etc.); external parasites (scabies, lice, ticks, etc.); blood parasites (Trypanosomiasis, Heartwater, Babesiosis, avian Spirochetosis, etc.);

- bacterial infections: contagious Pleuropneumonia of bovines and goats, Pasteurellosis of bovines and sheep, Anthrax and Blackleg, bovine Brucellosis and Tuberculosis, avian Salmonellosis;

- viral infections: Rinderpest and other plagues, Newcastle disease, Rabies, Blue Tongue, poxvirus with sheep, goat and camels, Lumpy Skin disease, Foot and Mouth disease

These diseases are subject to medical treatments (parasites), prophylaxis (Trypanosomiasis) or medical and sanitary prophylaxis (pests, Pleuropneumonia, clostridial infections, Salmonellosis).

Until recently, animal health affairs were exclusively managed by the government animal health services. Prophylaxis against the major animal diseases was mandatory and free. In 1979, with the founding of the central veterinary pharmacy (VETOPHAR, now LABOCEL), the livestock owner had to bear the costs. However, in spite of considerable rural extension, owners did not accept this change. So free vaccination was carried out until 1991. Today, vaccination is largely subsidized but still widely rejected by stock owners.

The current situation of animal health in Niger is characterized by the following situation:

Permanent pastures occupy approximately 60,000,000 ha in Niger and acquire their greatest extent in the Sahelian zone, the present Nigerien pastoral zone. The area of pastures is limited in the north by the semi-arid zone (100-mm isohyet) and to the south by the northern limit of agriculture, marked by the 350-mm isohyet. The location of the isohyets varies considerably from year to year. The ecological conditions make crop-raising a risky speculation, but are disposed towards livestock keeping, in spite of the variability of rainfall.

Water is a key factor for survival and development, ensuring the growth of natural pastures and the maintenance of the domestic herds: more than 3,000 water points were listed within the pastoral territory (1970). Thus, in Niger, pastoral management is basically a question of management of water, i.e. how to balance the size of the herd with the watering capacity of the water points and the family labour available to satisfy the watering of the animals in the minimum time and the best possible conditions.

The variability of rainfall involves unpredictability of fodder production, as much in space as for the food value of the pastures. Other factors also have to be taken into account, such as the influence of the selective grazing, natural and man-made fires and the way of exploitation of the pastures. The action of man can play a paramount role in the composition and structure of the vegetation by opening more or less the herbaceous soil protection, thus supporting or, on the contrary, limiting the growth of certain woody plants.

Owing to the variability of rainfall, annuals are often more competitive, especially on sandy soils. While the availability of water plays a paramount role in the productivity of the pastures in the northern part of the Sahelian zone, it appears that soil fertility acts as a limiting factor in the southern part where there is a better water supply. Fodder availability varies according to area and in general increases from north to south.

Forage in the Saharo-Sindian zone is often provided by perennials and supplemented by the irregular appearance of annuals induced by random precipitation. Only the transhumance of large herds of small ruminants and camels allows the rational exploitation of this sporadic vegetation (' acheb' of Touareg). Some Poaceae play a paramount role in livestock feed, like Panicum turgidum, Lasiurus scindicus, Chrysopogon plumulosus, Centropodia forskalii, Stipagrostis vulnerans and Stipagrostis acutiflora. Other annuals seem to be very significant: Schouwia thebaica (Brassicaceae), Citrullus colocynthis (Curcubitaceae) or Astragalus vogelii (Leguminoseae).

Unlike the Ahaggar area of Algeria where the pastures contain numerous plants rich in salt, the Aïr has none. Touaregs lead their herds to the south-west of the massif where groups of stockmen meet at the time of the 'salt cure' (salt licks for the herds) in Ighazer, north of InGall. The salt grounds of this area are covered with a very dense herbaceous layer in the rainy season: Ipomoea verticillata (Convolvulaceae), a dominant species in the herbaceous carpet, is regarded as best fodder for the goats, sheep and dromedaries.

The pastoral part of the Sahelian zone extends from West to East over 1,400 km through Niger and reaches a width of 250 km. According to the rainfall and geomorphology, the pastoral zone can be divided into three subzones: north, south and centre.
In the northern Sahelian zone (between the 100-mm isohyet in the south and the semi-desert of the north), on soils covered by sand (the majority of soils), trees are few and the commonest ones are Acacia raddiana, Commiphora africana, Leptadenia pyrotechnica and ubiquitous trees such Balanites aegyptiaca, Maerua crassifolia and Ziziphus mauritania. The latter two are relished by goats and camels throughout the year. On highly deteriorated soils, Calotropis procera pioneers, only eaten by goats and sheep when other plants are scarce. The herbaceous layer is dominated by perennial Poaceae , in particular by Panicum turgidum, a good fodder plant with stalks that stay green beyond the rain season, and various Stipagrostis spp. On the same soil Cornulaca monacantha and Cornulaca aucheri (Chenopodiaceae) can be found. These species constitute pastures that produce - under good conditions - approximately 400 kg of dry matter / ha. On soils with more clay and silt (depressions and valleys), the grass layer is mainly annual: Aristida hordeacea and Aristida funiculata are particularly frequent. The productivity of such pastures is slightly higher and can reach 500 kg of dry matter / ha.

On clayey soils an important plant formation is provided by Sorghum arundinaceum and Sporobolus helvolus which, in Ighazar (north of InGall), form immense pastures on salty land in the rainy season. The significant evaporation related to the endorheic water (internal drainage) has led to the accumulation of salts (sodium chloride and sulphate) in the surface layers of the soil. The very rich pasture on these grounds, is the cause of the gathering of many herds: it represents a true social rite at that time of the year, where the difficulties of the dry season disappear with the arrival of the rains. (There is a famous annual gathering of Touaregs and other Saharan nomads for a festival/reunion/market event at InGall, an oasis village two hours by road from Agadez. It is said to be one of the oldest and largest of such gatherings in the Sahara). This 'salt cure' (cure salée) is in fact a convalescence of the livestock after the long difficult months: a significant nutritious contribution of rock salt, a purgative action and intestinal worm cure for the animals.

The central Sahelian zone is between the 100-mm isohyet in the north and the 200 - 350-mm isohyet in the south. Annuals prevail in this subzone, not only due to a low rainfall but also because of fire and grazing. The open woody layer consists mainly of various Acacia and the species mentioned in the preceding zone. On sandy soils, the herbaceous layer is now more or less continuous and the commonest plants are Panicum turgidum, a perennial, and annuals such as Aristida mutabilis, Cenchrus biflorus, Eragrostis tremula, Brachiaria xantholeuca and Schoenefeldia gracilis. Other families are represented with annual plants such as Indigofera spp., Blepharis linariifolia, Gisekia pharnacioides, Borreria spp., Mollugo spp. and others. The productivity of these pastures reaches 1-1.5 ton of dry matter / ha. On soils with higher clay and silt content, mostly in depressions, the distribution of species is more heterogeneous and the herbaceous layer consists mainly of Schoenefeldia gracilis, Aristida funiculata and Cymbopogon schoenanthus. The latter formations are very seasonal, they appear with a relatively high productivity at the beginning of the rainy season but they dry very fast at its end.

The southern Sahelian subzone extends its border to the south until the northern limit of agriculture which corresponds to the 350-mm isohyet. It includes fallow land of former arable lands whose extension towards the north had once taken place in years of sufficient rainfall. Sandy soils on dunes or plateaux carry savanna formations of Mimosaceae and Combretaceae which show the traces of the degradations of the former extension of the agricultural land. Still, the grass layer contains a strong share of therophytes (annual plants that persevere in unfavourable climatic conditions as seeds) like Schoenefeldia gracilis, Aristida mutabilis and Cenchrus biflorus. The productivity of these pastures can exceed - depending on their floristic composition - two tons of dry matter per hectare. On soils with a higher clay and silt content, the grass cover does not differ very much from the one of the preceding central subzone. On hydromorphic soils - bordering Lake Chad - species such as Panicum repens, Panicum porphyrrhizos, Sporobolus spicatus and Sporobolus helvolus are dominant and on more sandy sites, Schoenefeldia gracilis and Echinochloa colona. They are excellent pastures that can be exploited during the whole year.

The whole sedentary part of the Sahelian zone is occupied by agriculture, on the clayey as well as on the more sandy soils. Grazing is thus possible only from November after harvest or on fallow land: in both cases, it mainly benefits domestic herds. The floristic composition of the fallow lands varies with their age and their productivity but does not exceed one ton of dry matter / ha. First, fallows are invaded by annual Poaceae, but later productivity improves with an increasing number of perennials: Eragrostis tremula, Aristida mutabilis, Aristida funiculata are thus replaced by Andropogon gayanus, Aristida sieberiana, Cymbopogon schoenathus and Hyperthelia dissoluta.

On ferralitic sandy soils of the savannas of the Sudanian zone, dominant plants of the grass layer are Panicum nigerense and Loudetia hordeiformis, on the slopes of the Dallol ('valley') Maouri Aristida mutabilis, Aristida sieberiana and on some isolated dunes Andropogon gayanus. These are found in company with Eragrostis tremula, Diheteropogon grandiflorus, Aristida stipoides, Cenchrus biflorus, Digitaria gayana and other grasses. On the more clayey soils of the wooded savannah along the Niger river, species such as Hyparrhenia involucrata, Hyparrhenia cyanescens, Brachiaria xantholeuca, Andropogon gayanus are found; on hydromorphic soils Hyperthelia dissoluta, Andropogon pseudapricus are frequent. On the plateaux, where we find the 'tiger bush' formation, the grass layer consists mainly of annuals such as Elionurus elegans, Andropogon pseudapricus, Andropogon fastigiatus, Pennisetum pedicellatum, Eragrostis tremula, Eragrostis pilosa, Dactyloctenium aegyptium, Aristida adscensionis, Schoenefeldia gracilis, Tripogon minimus, Brachiaria xantholeuca together with a few perennials that sometimes dominate: i.e. Loudetia simplex or Loxodera lermannii.

The Niger riverbed and temporary ponds are locally important pastures. 'Bourgoutières' are pastures that are temporarily flooded by the Niger river. In Niger, the most important 'bourgoutières' are close to the Malian border. These formations are subjected to two annual floods: a first one ('white water') between mid-September and mid-October, and a second one ('black water') between mid-February and mid-March. Different types of bourgoutières can be distinguished: The borgoutières with Brachiaria mutica, the bourgoutières with Echinochloa stagnina (the 'bourgou'), and the bourgoutières with Oryza longistaminata and Echinocloa stagnina. As the 'white water' withdraws (mid-October), the animals enter the bourgoutières with Brachiaria mutica and leave them only in December. The dry stalks are then burned to make them resprout. The livestock then returns to these sites as soon as the 'black water' withdraws in March. The bourgoutière with Echinochloa stagnina is grazed later, in April, and the herds remain there until the arrival of the white water. The productivity of the bourgoutières is in general high: for the formations with Echinochloa stagnina, it varies from 6 to 17 tons of dry matter / ha (in Mali, it can reach 30-40 tons). Unfortunately, under the effect of water deficit due to climatic changes, overgrazing, the exploitation of grass tufts (the grass is used for fattening domestic animals which are then sold on the markets), extensive rice growing and cultivation of the river banks, one observes a clear regression of the bourgoutières and an evolution of their floristic composition which has led to a drastic reduction in the fodder resources.

The fallow lands of the Sudanian area are, as per the fallow land of the Sahelian area, complementary fodder resources. At the beginning, the fallow land is dominated by Eragrostis tremula, associated with Ctenium elegans, Loudetia hordeiformis and Schizachyrium exile. They are later partly replaced by species such as Pennisetum pedicellatum and Diheteropogon hagerupii. The productivity of these fallows can be up to 4 tonnes of dry matter / ha.

Table 3. Productivity and Carrying Capacity of Different Pastoral Zones

Fodder. For farmers, weeds cause more harm than good if not controlled, and weed control causes a labour peak in crop production. Howver, livestock plays a key role in the traditional household economy and various weeds are traded as feed, food or construction material. Especially in mixed-farming systems, in the rainy season, feed demands of animals can be satisfied by fresh grasses and herbaceous plants. At the onset of the dry season, animals prefer millet leaves and weeds to millet stalks. During the dry season, feed is mostly limited to the scarce biomass from natural pastures and the decomposition of stover. At rural markets, fodder weeds such as Alysicarpus ovalifolius, Zornia glochidiata (both Leguminoseae), Commelina forskalaei (Commelinaceae) or Eragrostis tremula (Poacea) are sold. Laboratory analyses of these weeds revealed a crude protein content twice that of low quality roughages such as millet stover.

Livestock keepers show a preference for different weeds, such as Eragrostis tremula, Commelina forskalaei, Cenchrus biflorus (Poaceae) and Merremia tridentata (Convulvaceae), a perennial creeper. The grass Eragrostis tremula can be cut several times during the rainy period and is eaten by all animals. It can be ensiled (another use is broom production). Commelina forskalaei is a good fodder and suited for the preparation of silage. Under wet weeding conditions it regenerates fast. Both Eragrostis tremula and Commelina forskalaei emerge after the first weeding. Cenchrus biflorus is difficult to weed and rather persistent under humid conditions. It is grazed by most animals in its juvenile state and standing hay after the fall of the prickly spikelets. The fermentation process during ensiling softens the bristles which can subsequently be eaten by animals without harm. It is a reliable feed, since it persists until the next rainy period. Women gather Cenchrus biflorus seeds to prepare a porridge after cracking the seeds. Merremia tridentata is appreciated by all animals and remains on the grassland for about four months after the rainy period. It is well suited for preparing hay and silage. Further livestock diet improvement is attributed to Mitracarpus scaber (Rubiaceae) and Ceratotheca sesamoides (Pedaliaceae). They are not suitable for haymaking but for silage. The leaves of Ceratotheca sesamoides are used as a vegetable. Mitracarpus scaber remains green for about three months after the rainy season.

In the Sahelian zone, most grasses are well consumed by animals; some are preferred to others. Generally, those that grow on humid ground have larger leaves and are more tender (e.g. Echinocloa colona, Panicum laetum). Some grasses are more or less 'ligneous' and are disdained, especially when dry (e.g. Aristida hordeacea, Elionurus elegans). In the north of the Sahelian zone, grasses such as Panicum turgidum, Stipagrostis vulnerans or Lasiurus scindicus are relished by all animals and sometimes taken as hay to feed the animals of the moving caravans.

The degree of acceptance of weeds by animals depends on many factors such as season and food scarcity. Under conditions of severe food shortage, even toxic plants are eaten, especially by goats. While the biomass increases with the age of the plants, the fodder value decreases sharply after stem elongation. During the dry season, the fodder value of straw is low and has to be compensated by the consumption of leaves and fruits of trees.
Generally, the mineral content of weeds depends on their growth stage. In most cases in a tropical environment, grasses are low in sodium. During the stage of sprouting, they are rich in phosphorus, but its content and the calcium content decrease towards the end of the growth cycle. The content of cobalt and selenium is usually satisfactory, but zinc and copper are often deficient in straw, whereas a lack of iron is rarely found in tropical environments.

Table 4. Consumption of Fodder Trees

x = eaten , r = limited palatability, ? = no information, n = no consumption

Table 5. The Nutritional Value of Fodder