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4. SELECTED STRADDLING FISH STOCKS


The list of straddling stocks in FAO (1994) was taken as a starting point for this review. Enquiries were sent to Regional Fisheries Bodies soliciting regional knowledge to refine the lists. Information was received for the Northeast and the Southeast Atlantic. For the northwest and Southwest Atlantic, and the Northeast, eastern central and Southeast Pacific, staff of the FAO Fisheries Department or its consultants applied their own informed judgements. For the western and eastern central Atlantic, the Southwest Pacific and for the Indian Oceans, catches by country (within statistical areas) were examined to determine which species were being reported by non-coastal States which were presumed to be fishing on the high seas. This information was tempered by knowledge of situations where distant-water fishing countries have access agreements to EEZs, particularly when the species in the reported catch were not known to be in commercial abundance on the high seas. Using this approach, a refined list of species (by FAO statistical area) likely to be fished as straddling stock and high seas fish stocks was prepared. This included 129 species/stocks and a brief description of the main, straddling stocks by major area as given below, and in Section 5 the same is done for the high seas fish stocks.

An examination of the continental shelves, defined for the purpose of this review by the 200-m depth contour, and the 200 nm contour from the coastline which generally corresponds to EEZs (Figure 45), suggests that there are several areas where stocks of neritic fishes and in particular of demersal fishes associated with the continental shelves could be straddling: a small area north of Russia and the United States of America in the North Pacific, a small area off Argentina in the Southeast Atlantic, the nose and tail of the Newfoundland Grand Bank off Canada in the northwest Atlantic, an area between Norway and Russia in the Northeast Atlantic (the so-called Loop Hole), an area off Mauritius and the Seychelles in the western Indian Ocean, and another area off the Russian EEZ in the northwest Pacific Ocean. Other known areas of straddling stocks are the "Donut Hole" in the middle of the Bering Sea in the Northeast Pacific, the "Peanut Hole" in the Sea of Hokhotsk in the northwest Pacific, the Challenger Plateau west of New Zealand and the South Tasmanian Rise in the eastern Indian Ocean. This implies that the United States of America, Russia, Argentina, Canada, Norway, Australia and New Zealand are the coastal States most likely affected or potentially affected by demersal straddling stocks problems. Meltzer (2005) comes to a similar conclusion about areas with the potential of straddling stocks, highlighting in addition the eastern central Atlantic (FAO area 34) off West Africa, the Southwest Atlantic (FAO area 41) and off almost the entire east coast of South America.

FIGURE 45 Continental shelf areas within and beyond the 200 nautical mile limit off the coast

The main species that constitute straddling stocks are generally well studied (e.g. cod, pollock, flounders) compared to several highly migratory species, particularly the non-tunas. Such information is readily available from various published sources of information or reports or web pages from Regional Fishery Bodies (RFB). Therefore, this document does not review the biology, and life history and migratory behaviour of these species. As indicated before, there is no global database distinguishing between catches taken from (entirely) EEZ stocks from those taken from straddling stocks, or to distinguish catches of straddling stocks within and outside EEZs. Therefore, the catch data presented later in this section is only provided as an indication of the relative importance of the species, with no indication of what portion of it comes from straddling stock(s), or from areas within or beyond the EEZ. The only graphs of catches presented in this section are for the Southern Ocean, where we report on all of the species fished in the CCAMLR convention area as if they were straddling stocks.

4.1. Pacific Ocean

4.1.1 Northwest Pacific

Straddling stocks in the northwest Pacific include Alaska (Walleye) pollock (Theragra chalcogramma), flying squid (Ommastrephes bartrami), Boreal clubhook squid (Onychoteuthys borealjaponica), Boreopacific armhook squid (Gonatopsis borealis), Pacific Ocean perch (Sebastes alutus), armourhead (Pseudopentaceros spp.) and the alfonsino (Beryx splendens). Based on FAO (2005a), the pollock is considered fully exploited while the squids vary from moderately to fully exploited, and in some cases recovering. Based on reported landings, the Pacific Ocean perch is considered depleted, while the state of armourhead and alfonsinos is not known.

4.1.2 Northeast Pacific

Straddling stocks in the Northeast Pacific include jack mackerel (Trachurus picturatus symmetricus) and Alaska (Walleye) pollock (Theragra chalcogramma). The jack mackerel is moderately exploited and the Alaska pollock is fully exploited.

4.1.3 Western central Pacific

There is no information on straddling stocks in the western central Pacific.

4.1.4 Eastern central Pacific

According to FAO (2005a) the straddling stocks of jumbo flying squid (Dosidicus gigas) are moderately to fully exploited in the eastern central Pacific, horse mackerel (Trachurus spp.) is underexploited, and chub mackerel (Scomber japonicus) is moderately exploited.

4.1.5 Southwest Pacific

There exist two types of straddling fish stocks in the Southwest Pacific. Stocks belonging to the more common type are associated with continental shelves while stocks belonging to the less common type are associated with small islands with limited shelves whose fishery depend on oceanic resources found both within and outside their EEZs. Species with straddling stocks associated with continental shelves include orange roughy (Hoplostethus atlanticus), oreo dories (Allocyttus verrucosus, A. niger, Neocyttus rhomboidalis, Pseudocyttus maculatus) and hoki (Macruronus novaezealandiae). Straddling oceanic resources include the narrow-barred Spanish mackerel (Scomberomorus commerson), oceanic squids, and flying fish.

Based on FAO (2005a), orange roughy, oreo dories and hoki are fully exploited to overexploited. The Spanish mackerel are moderately exploited, oceanic squid are moderately exploited and flying fish are fully exploited.

4.1.6 Southeast Pacific

Straddling stocks in the Southeast Pacific include jumbo squid (Dosidicus gigas) and Chilean jack mackerel (Trachurus picturatus murphyi). Chub mackerel (Scomber japonicus) is also found beyond the EEZ but the catches are small. The Chilean jack mackerel is fully or overexploited while the jumbo flying squid is moderately exploited based on FAO (2005a). Chub mackerel is reported as moderately to fully exploited.

4.2 Atlantic Ocean

4.2.1 Northwest Atlantic

Straddling stocks in the northwest Atlantic include cod (Gadus morhua), American plaice (Hypoglossoides platessoides), redfish (Sebastes marinus), witch flounder (Glyptocephalus cynoglossus), Atlantic halibut (Hippoglossus hippoglossus), black halibut (Reinhardtius hippoglossoides), yellowtail flounder (Pleuronectes ferruginaeus), grenadiers (Macrouridae), capelin (Mallotus villosus) and shrimp (Pandalus borealis).

Based on assessments by the Northwest Atlantic Fisheries Organization (NAFO) where stocks are not assessed every year (a convenient summary is found at http://www.nafo.ca/science/advice/nafo-stocks.html), cod, American plaice, redfish, witch flounder, and Atlantic halibut are depleted; black halibut are overexploited; yellowtail flounder and shrimp are fully exploited; capelin are underexploited, and the status of grenadiers is unknown. It is noted that stocks of some of the species on the Flemish Cap (NAFO Division 3M), such as cod and redfish, may be separate from EEZ stocks, and as such, may be high seas fish stocks, rather than straddling stocks.

4.2.2 Northeast Atlantic

The main "traditional" straddling stocks in the Northeast Atlantic are: blue whiting (Micromesistius poutassou), oceanic redfish (Sebastes mentella), cod (Gadus morhua), haddock (Melanogrammus aeglefinus), black halibut (Reinhardtius hippoglossoides), Norwegian spring-spawning herring (Clupea harengus), mackerel (Scomber scombrus) and horse mackerel (Trachurus trachurus).

In addition to these, the Northeast Atlantic Fisheries Commission (NEAFC, http://www.neafc.org) advises in its reply to FAO's request for input, that most deepwater species for which fisheries have recently developed should also be considered as being straddling. These species are: Baird's smoothhead (Alepocehalus bairdii), Risso's smoothhead (Alepocephalus rostratus), blue antimora or blue hake (Antimora rostrata), black scabbardfish (Aphanopus carbo), Iceland catshark (Apristuris spp.), greater silver smelt (Argentina silus), alfonsinos (Beryx spp.), tusk (Brosme brosme), gulper shark (Centrophorus granulosus), leafscale gulper shark (Centrophorus squamosus), black dogfish (Centroscyllium fabricii), Portuguese dogfish (Centroscymnus coelolepis), longnose velvet dogfish (Centroscymnus crepidater), deep-water red crab (Chacon (Geyron) affinis), rabbit fish (Rattail) (Chimaera monstrosa), frilled shark (Chlamydoselachus anguineus), conger eel (Conger conger), roundnose grenadier (Coryphaenoides rupestris), kitefin shark (Dalatias licha), birdbeak dogfish (Deania calceus), black (deep water) cardinal fish (Epigonus telescopus), greater lanternshark (Etmopterus princes), velvet belly (Etmopterus spinax), blackmouth dogfish (Galeus melastomus), mouse catshark (Galeus murinus), bluemouth (blue mouth redfish) (Helicolenus dactylopterus), blondnose six-gilled shark (Hexanchus griseus), orange roughy (Hoplostethus atlanticus), Mediterranean slimehead (Hoplostethus mediterraneus), large-eyed rabbit fish (ratfish) (Hydrolagus mirabilis), silver scabbard fish (cutless fish) (Lepidopus caudatus), greater eelpout (Lycodes esmarkii), roughhead grenadier (rough rattail) (Marcrourus berglax), blue ling (Molva dypterigia), ling (Molva molva), common mora (Mora moro), sailfin roughshark (sharpback shark) (Oxynotus paradoxus), red (blackspot) seabream (Pagellus bogaraveo), forkbeards (Phycis spp.), wreckfish (Polyprion americanus), round skate (Raja fyllae), Arctic skate (Raja hyperborea), Norwegian skate (Raja nidarosiensus), straightnose rabbitfish (Rhinochimaera atlantica), knifetooth dogfish (Scymnodon ringens), small redfish (Norway haddock) (Sebastes viviparous), Greenland shark (Somniosus microcephalus), spiny (deep sea) Atlantic thornyhead (Trachyscorpia cristulata).

Fisheries for deep-water species have developed rapidly since 1990 in the Northeast Atlantic as a result of management limitations and reduced resource availability of traditional species. The International Council for the Exploration of the Sea (ICES,2005) has recently provided advice for the management of these fisheries. Although the state of exploitation of most species/stocks cannot be assessed with respect to standard criteria, the overall evaluation provided by ICES is that those fisheries are currently not sustainable. Based on ICES advice in 2005, none of the traditional straddling stocks are under or moderately exploited, herring and oceanic redfish are fully exploited, blue whiting, cod, haddock, black halibut and mackerel are overexploited, while the state of horse mackerel is uncertain.

4.2.3 Eastern central Atlantic

The analysis of catches by non-coastal States described in the introduction to the chapter on straddling stocks identified catches of common cuttlefish, marine fishes nei, Octopuses etc. nei, red porgy, West African goatfish, common sole, cuttlefish, bobtail squids nei, European hake, Natantian decapods nei, croakers, drums nei, tonguefish, chub mackerel, European pilchard, jack and horse mackerel nei, alfonsinos, flatfishes nei and Senegalese hake from countries that have fishing agreements with coastal States. All of these species are distributed within EEZs and, as generally understood among the CECAF working groups, it is concluded that there are no significant fisheries for straddling stocks outside of EEZs at present in the eastern central Atlantic.

4.2.4 Western central Atlantic

The analysis of catches by non-coastal States was also performed for the western central Atlantic. It identified catches of a mixture of coastal and oceanic species in general categories such as: sharks, rays, skates, etc. nei; croakers, drums nei; hairtails, scabbardfishes nei; marine fishes nei, and Natantian decapods nei, which suggest that these catches were probably made within EEZs under fishing agreements with coastal States. As for the eastern central Atlantic, it was concluded that there are no significant fisheries for straddling stocks outside EEZs at present in the western central Atlantic.

4.2.5 Southwest Atlantic

Straddling stocks in the in the Southwest Atlantic include short-fin squid (Illex argentinus), common squid (Loligo spp.), seven star flying squid (Martialia hyadesi), the Argentine hake (Merluccius hubbsi) and southern hake (M. polylepis), the southern blue whiting (Micromesistius australis), the pink cusk eel (Genypterus blacodes), the Patagonian toothfish (Dissostichus eleginoides), the tadpole mora (Salilota australis), the Patagonian grenadier (Macruronus magellanicus), the grenadier (Macrourus whitsoni), the Antarctic cod (Notothenia rossii), rockcods (Notothenia spp.) and various species of sharks and rays.

Based on FAO (2005a) the state of exploitation of the stocks of short-fin squid and common squid is fully exploited, while the state of the seven star flying squid will be unknown. However, given its life history pattern, relatively wide distribution and low and occasional nature of the catches, it is unlikely that it will be fully or overexploited. The state of Argentine hake (mostly found and caught within the EEZs) are reported as overexploited or depleted, while the southern hake is considered fully exploited. The southern blue whiting is fully to overexploited, and the pink cust eel and the Patagonian toothfish are moderately to fully exploited, although FAO (2005a) notes that "there is a general perception that Patagonian toothfish is in a much critical situation due to high non-reported catches in international waters". The Patagonian grenadier is moderately exploited while the state of the tadpole mora, rockcods and the shark and rays are not known.

4.2.6 Southeast Atlantic

In reply to FAO's request to RFB's, the Southeast Atlantic Fisheries Organization (SEAFO, http://www.seafo.org/) identified the following species as straddling: alfonsinos (family Bercycidae), orange roughy, horse mackerel (Trachurus spp.), lanternfish (family Myctophidae), mackerel (Scomber spp.), skates (family Rajidae), sharks (Order Selachomorpha), armourhead (Pseudopentaceros spp.), cardinal fish (Epigonus spp.), deep-sea red crab (Chaceon maritae), octopus (family Octopodidae), squids (family Loliginidae), and wreckfish (Polyprion americanus). The state of exploitation is unknown for all of the species except for horse mackerel which are classified as fully exploited.

4.3 Indian Ocean

The narrow continental margins around the Indian Ocean result in few straddling stocks. The transboundary and straddling stocks that do exist (e.g. deep-water snapper, deep-water shrimp) are minor and are not fished to any extent: there are no high seas fisheries for these resources. The topography of the Mascarene Ridge makes this area suitable for straddling stocks as its shallow waters extends beyond the EEZ onto the high seas. Here, an important example is the Saya de Malha Bank where stocks of dame berri (Lethrinus mahsena) and capitaine (L. nebulosa) are fished within, but mainly beyond the Mauritian EEZ.

4.4 Southern Ocean

The Southern Ocean is considered to be delimited by the Antarctic Convergence where cold Antarctic waters meet warmer waters of the Atlantic, Pacific and Indian Oceans to the north. The location of the Antarctic Convergence varies over time, but it is in the vicinity of 60 °S. The Antarctic convergence is generally considered to form the boundary of the Southern Ocean ecosystem, with relatively few species passing through it (the great whales are a notable exception).

There is a unique legal situation for the Southern Ocean (FAO statistical areas 48, 58 and 88) which is within the jurisdiction of the Commission for the Conservation of Antarctic Marine Living Resources (CCAMLR, http://www.ccamlr.org/default.htm). Claims of sovereignty over the Antarctic Continent or its continental shelf have been put aside under provisions of the Antarctic Treaty, which entered into force in 1961 (http://www-old.aad.gov.au/information/treaty/treaty.asp). However, several countries have established EEZs within the Southern Ocean area of CCAMLR off the coasts of their island territories in that area, in addition to the EEZs extending from the tip of South America. Given the unique situation of the Southern Ocean with respect to territorial and jurisdictional claims, this review reports on all of the species fished in the CCAMLR convention area as if they were straddling fish stocks or high seas fish stocks. There are no fisheries for highly migratory species (as defined by Annex 1 of UNCLOS) in the Southern Ocean at present. Cetaceans or whales are listed as highly migratory species in Annex 1 of UNCLOS and, historically, the Southern Ocean was a major area for whaling. However, the International Whaling Commission currently maintains a moratorium on commercial whaling.

Prior to the mid-1960s, only whale catches were reported to FAO from the Southern Ocean. Since then, the fisheries have targeted various species including marbled rockcod, mackerel icefish, humped rockcod, south Georgian icefish, Patagonian and Antarctic toothfish and Antarctic krill. Reported catches exceeded 600 000 tonnes in the early-1980s, but since the early-1990s, they have been relatively stable around 100 000 tonnes per year, albeit with a tendency to increase. During 1990 - 2004 (Figures 46 and 47), the catches have been dominated by Antarctic krill (86 percent), Patagonian toothfish (6 percent), a lanternfish (Electrona carlsbergi), the subantarctic electron (5 percent) and the mackerel icefish at less than 2 percent. More than 50 species are reported in the remaining 1 percent of the total catches. Although Antarctic toothfish does not show up as an important species in the catch statistics, it is a concern within the convention area of CCAMLR because of IUU fishing during the late-1990s and early-2000s.

FIGURE 46
Catches by species in the Southern Ocean as reported to FAO in percentage cumulative, 1990 - 2004


FIGURE 47
Catches reported to FAO from the Southern Ocean

Information on the state of exploitation of Southern Ocean resources was provided by the CCAMLR Secretariat. Antarctic krill (Euphausia superba) is considered underexploited in FAO areas 48 and 58, while the lanternfish subantartic electron (Electrona carlsbergi), the sevenstar flying squid (Martialia hyadesi), the red stone crab (Paralomis spinosissima) and the globose king crab (P. formosa) in FAO area 48 are also considered underexploited. Patagonian toothfish (Dissostichus eleginoides) is considered overfished in parts of FAO area 58 and fully exploited in area 48 and other parts of area 58. Mackerel icefish (Champsocephalus gunnari) is fully exploited in both areas 48 and 58 while the state of marbled rockcod (Notothenia rossii), on which the fishery is closed, is uncertain. No species are considered moderately exploited in the Southern Ocean.

4.5 Mediterranean Sea and Black Sea

No EEZs have been implemented in the Mediterranean Sea. Therefore, all fish stocks exploited within and beyond the 12 nm of territorial sea will correspond to the definition of a straddling stock. Because of the shelf configuration, fewer bottom resources are likely to be straddling except in the Gulf of Sirte, off Tunisia and Libya. Most pelagic stocks, however, including sardine, jack mackerel and mackerel stocks, are most probably straddling. The General Fisheries Commission for the Mediterranean (GFCM) uses the concept of "shared" stocks to identify those exploited by two or more countries on the high seas and only by the riparian countries in territorial waters. Such stocks in the Mediterranean include hake (Merluccius merluccius) in the Gulf of Lions, deep-sea shrimps, the blue and red shrimp (Aristeus antennatus) and the giant red shrimp (Aristaeomorpha foliacea), sardines (Sardina pilchardus) in the Sea of Alboran and Adriatic Sea and anchovy (Engraulis encrasicolus) in the Gulf of Lions and the Adriatic Sea. Based on FAO (2005a), the state of exploitation of giant red shrimp is not known, the rose shrimp is fully exploited and the hake is overexploited. The state of sardines and anchovies ranges from underexploited to overexploited depending on the zone. It should be noted that fishing with towed gears beyond 1 000 m depth is forbidden by GFCM. As the riparian States of the Black Sea have implemented EEZs till the median line, there are no international waters in the Black Sea and, by definition, no straddling stocks.

4.6 State of selected straddling stocks

The state of exploitation of the main species-area combinations that include straddling fish stock is summarized in Table 4. The state of exploitation is not known for 20 percent of the species/stock categories used. Overall, taking those for which information is available, it is found that 4 percent of the straddling stocks are underexploited, 12 percent are moderately exploited, 19 percent are fully exploited, 58 percent are overexploited, 6 percent are depleted and 1 percent are recovering. Based on ICES (2005) advice to NEAFC that fisheries for deep-water species (which NEAFC categorized as straddling stocks), were not sustainable, they have been classified herein as overfished. This may overstate the percentage of stocks that are overexploited, because some stocks might not yet be overexploited and management interventions may prevent them from becoming overexploited. Alternatively, their state might have been classified as unknown, given ICES' reservations about data quality and its ability to assess the state of exploitation, but in light of ICES' concerns about sustainability, it seems prudent to classify these stocks as overexploited, unless demonstrated otherwise.

4% of the straddling fish stocks are underexploited, 12% moderately exploited, 19% fully exploited, 58% overexploited, 6% depleted and 1% recovering

TABLE 4
Summary of the state of exploitation of the main species-area combinations that may contain straddling fish stocks1

Species/stocks

Major ocean area

Catches (tonnes)2

State of exploitation3

2000

2001

2002

2003

2004

Sharks
(Selachimorpha)

SE Atlantic

N/A

N/A

N/A

N/A

N/A

N

Sharks, rays
(Elasmobranchii)

SW Atlantic

17 607

14 705

13 736

14 141

12 297

N

Frilled shark
(Chlamydoselachus anguineus)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Bluntnose sixgill shark
(Hexanchus griseus)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Blackmouth dogfish
(Galeus melastomus)

NE Atlantic

45

34

280

218

158

N-O4

Mouse catshark
(Galeus murinus)

NE Atlantic

45

34

280

218

158

N-O4

Deep-water catsharks
(Apristurus spp.)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Iceland catshark
(Apristurus laurussonii)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N

Greenland shark
(Somniosus microcephalus)

NE Atlantic

45

58

56

61

66

N-O4

Gulper shark
(Centrophorus granulosus)

NE Atlantic

141

248

401

919

672

N-O4

Leafscale gulper shark
(Centrophorus squamosus)

NE Atlantic

1 937

1 895

3 042

2 947

2 556

N-O4

Velvet belly
(Etmopterus spinax)

NE Atlantic




5

10

N-O4

Great lanternshark
(Etmopterus princeps)

NE Atlantic




5

10

N-O4

Birdbeak dogfish
(Deania calceus)

NE Atlantic

18

51

102

140

323

N-O4

Portuguese dogfish
(Centroscymnus coelolepis)

NE Atlantic

1 861

3 225

3 687

4 232

4 021

N-O4

Longnose velvet dogfish
(Centroscymnus crepidater)

NE Atlantic

1

3

16

509

301

N-O4

Knifetooth dogfish
(Scymnodon ringens)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Kitefin shark
(Dalatias licha)

NE Atlantic

311

189

40

735

603

N-O4

Black dogfish
(Centroscyllium fabricii)

NE Atlantic

271

271

27

53

56

N-O4

Sailfin roughshark
[Sharpback shark]
(Oxynotus paradoxus)

NE Atlantic



1


1

N-O4

Rays and skates nei
(family Rajidae)

SE Atlantic

N/A

N/A

N/A

N/A

N/A

N

Round ray
(Raja fyllae)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Arctic skate
(Raja hyperborea)

NE Atlantic




5


N-O4

Norwegian skate
(Raja nidarosiensus)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Rabbit fish [Rattail]
(Chimaera monstrosa)

NE Atlantic

15

122

69

169

617

N-O4

Large-eyed rabbitfish
[Ratfish]
(Hydrolagus mirabilis)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Straightnose rabbitfish
(Rhinochimaera atlantica)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Atlantic Herring,
Norwegian spring spawning
(Clupea harengus)

NE Atlantic (Atlanto-Scandian)

2 103 709

1 645 085

1 614 754

1 661 405

1 755 260

F

Sardines
(Sardina pilchardus)

Mediterranean

215 932

199 927

189 128

179 178

187 442

U-O

European anchovy
(Engraulis encrasicolus)

Mediterranean

391 757

438 381

481 327

400 636

455 062

U-O

Capelin
(Mallotus villosus)

NW Atlantic

21 374

19 751

13 646

22 455

36 477

U

Greater argentine
(Argentina silus)

NE Atlantic

28 533

48 731

37 033

21 395

32 865

N-O4

Baird's slickhead
(Alepocehalus bairdii)

NE Atlantic

12

616

259

9 663

7 492

N-O4

Risso's smooth-head
(Alepocephalus rostratus)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Lanternfish
(family Myctophidae)

SE Atlantic

N/A

N/A

N/A

N/A

N/A

N

Greater eelpout
(Lycodes esmarkii)

NE Atlantic

28

37

43

30

49

N-O4

European conger
(Conger conger)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Flying fishes nei
(Exocoetidae)

SW Pacific

N/A

N/A

N/A

N/A

N/A

M

Common mora
(Mora moro)

NE Atlantic

130

351

109

226

6

N-O4

Blue hake
(Antimora rostrata)

NE Atlantic

0

0

0

0

0

N-O4

Tadpole codling
(Salilota australis)

SW Atlantic

15 555

8 151

4 648

7 595

6 299

N

Tusk
(Brosme brosme)

NE Atlantic

31 246

27 019

25 815

20 760

19 039

N-O4

Atlantic cod
(Gadus morhua)

NE Atlantic

877 150

884 785

847 874

807 870

859 919

O

Atlantic cod
(Gadus morhua)

NW Atlantic

63 201

60 074

55 337

41 132

39 649

D

Ling
(Molva molva)

NE Atlantic

43 320

37 341

41 552

40 410

35 380

N-O4

Blue ling
(Molva dypterigia)

NE Atlantic

16 146

18 669

12 538

11 173

7 785

N-O4

Forkbeards
(Phycis spp.)

NE Atlantic

496

475

463

1 081

855

N-O4

Haddock
(Melanogrammus aeglefinus)

NE Atlantic

196 111

208 290

247 470

258 707

299 953

O

Alaska pollock [walleye]
(Theragra chalcogramma)

NE Pacific

1 183 482

1 443 917

1 519 122

1 530 299

1 522 860

F

Alaska pollock [walleye]
(Theragra chalcogramma)

NW Pacific

1 754 748

1 700 548

1 135 732

1 357 663

1 169 079

F

Blue whiting
(Micromesistius poutassou)

NE Atlantic

1 445 788

1 793 954

1 557 688

2 373 128

2 418 198

O

Southern blue whiting
(Micromesistius australis)

SW Atlantic

84 321

78 736

58 843

60 546

76 596

F-O

Hakes
(Merluccius spp.)

SW Atlantic


354

186


1 434

F-O-D

European hake
(Merluccius merluccius)

Mediterranean

68 897

52 408

53 950

55 037

62 951

O

Patagonian grenadier
(Macruronus magellanicus)

SW Atlantic

142 676

136 366

126 436

122 844

145 224

M

Hoki
(Macruronousnovaezelandiae)

SW Pacific

274 615

247 841

215 302

209 414

154 532

M-F

Grenadiers
(Macrourus spp.)

SW Atlantic

10 503

3 209

6 052

8 253

5 265

N

Roughhead grenadier
(Macrourus berglax)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Roughhead grenadier
(Macrourus berglax)

NW Atlantic

N/A

N/A

N/A

N/A

N/A

N

Roundnose grenadier
(Coryphaenoides rupestris)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Pink cusk-eel
(Genypterus blacodes)

SW Atlantic

17 521

22 702

19 712

16 450

19 293

M-F

Alfonsino
(Beryx spp.)

NW Pacific

18

14

12

44

1

N

Alfonsino
(Beryx spp.)

NE Atlantic

139

130

272

949

1 117

N-O4

Alfonsino
(Beryx spp.)

SE Atlantic

302

318

236

229

324

N

Mediterranean slimehead (Hoplostethus mediterraneus)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Orange roughy
(Hoplostethus atlanticus)

NE Atlantic

1 467

4 672

5 438

872

1 240

N-O4

Orange roughy
(Hoplostethus atlanticus)

SW Pacific

18 007

14 303

18 515

18 346

18 157

F-O4

Oreodories
(family Oreosomatidae)

SE Atlantic

10

54

335

331

497

N

Oreodories
(family Oreosomatidae)

SW Pacific

22 775

24 165

17 635

15 263

19 787

F-O

Wreckfish
(Polyprion americanus)

NE Atlantic

441

414

432

521

358

N-O4

Wreckfish
(Polyprion americanus)

SE Atlantic

8


2

6

1

N

Jack and horse mackerels nei
(Trachurus spp.)

EC Pacific

N/A

N/A

N/A

N/A

N/A

U

Jack and horse mackerels nei
(Trachurus spp.)

NE Atlantic

36 989

39 824

33 429

32 821

35 871

N

Jack mackerel
(Trachurus spp)

NE Pacific

N/A

N/A

N/A

N/A

N/A

M

Jack and horse mackerels nei
(Trachurus spp.)

SE Atlantic




1 941

234

F

Chilean jack mackerel
(Trachurus murphyi)

SE Pacific

1 540 494

2 508 834

1 750 078

1 736 048

1 778 777

F-O

Blackspot (=red) seabream
(Pagellus bogaraveo)

NE Atlantic

3 428

3 144

3 711

3 531

3 981

N-O4

Pelagic armourhead
(Pseudopentaceros richardsoni)

NW Pacific

N/A

N/A

N/A

N/A

N/A

N

Armourhead
(Pseudopentaceros spp.)

SE Atlantic

0

0

0

0

0

N

Antarctic rockcods, noties neii
(Nototheniidae)

SW Atlantic

N/A

N/A

N/A

N/A

N/A

N

Antarctic toothfish
(Dissostichus mawsoni)

Southern Ocean

751

626

1 354

2 029

2 584

F-O

Patagonian toothfish
(Dissostichus eleginoides)

Southern Ocean

16 911

13 179

13 989

16 479

11 182

F-O

Patagonian toothfish
(Dissostichus eleginoides)

SW Atlantic

11 128

13 823

12 488

8 871

6 671

M-F

Marble rockcod
(Notothenia rossii)

Southern Ocean

0

0

0

0

0

N

Humped rockcod
(Notothenia gibberifrons)

Southern Ocean

1

2

1



N

Antarctic rockcods
(Trematomus spp.)

SW Atlantic

0

0

0

0

0

N

Mackerel icefish
(Champsocephalus gunnari)

Southern Ocean

4 251

2 096

3 532

4 331

2 762

F-O

South Georgia icefish
(Pseudochaenichthys georgianus)

Southern Ocean


6

6

5

1

N

Cardinal fishes nei.
(Epigonus spp.)

SE Atlantic

N/A

N/A

N/A

N/A

N/A

N

Black cardinal fish
(Epigonus telescopus)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Mackerel,
(Scomber spp.)

SE Atlantic

N/A

N/A

N/A

N/A

N/A

N

Atlantic mackerel
(Scomber scombrus)

NE Atlantic

654 829

660 188

684 829

601 685

587 072

O

Chub mackerel
(Scomber japonicus)

SE Pacific

254 524

627 466

393 142

699 714

691 978

M-F

Spanish mackerel
(Scomberomorus sp.)

SW Pacific

1 601

2 079

1 327

897

647

M

Atlantic Spanish mackerel
(Scomberomorus maculatus)

EC Pacific

N/A

N/A

N/A

N/A

N/A

M

Silver scabbardfish [Cutlass fish]
(Lepidopus caudatus)

NE Atlantic

104

361

1 970

1 000

806

N-O4

Black scabbardfish
(Aphanopus carbo)

NE Atlantic

8 184

10 821

11 377

8 215

8 092

N-O4

Norway redfish
(Sebastes viviparus)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Atlantic redfish
(Sebastes spp.)

NW Atlantic

47 160

49 629

56 566

64 643

31 905

D

Pacific ocean perch
(Sebastes alutus)

NW Pacific

1 475

1 461

1 948

1 451

1 378

D

Beaked redfish
(Sebastes mentella)

NE Atlantic

75 506

98 534

92 344

9 469

83 968

F

Blackbelly rosefish
(Helicolenus dactylopterus)

NE Atlantic

743

624

453

488

504

N-O4

Atlantic thornyhead
(Trachyscorpia cristulata)

NE Atlantic




41

42

N-O4

Atlantic halibut
(Hippoglossus hippoglossus)

NW Atlantic

1 372

2 367

1 893

2 244

2 056

D

Greenland halibut
(Reinhardtius hippoglossoides)

NW Atlantic

64 583

61 431

64 482

71 859

56 893

O

Greenland halibut
(Reinhardtius hippoglossoides)

NE Atlantic

43 108

47 190

43 393

48 050

53 013

N-O4

Greenland halibut
(Rheinhardtius hippoglossoides)

NW Atlantic

64 583

61 431

64 482

71 859

56 893

O

Which flounder
(Glyptocephalus cynoglossus)

NW Atlantic

6 447

7 277

7 077

7 114

6 178

D

American plaice
(Hippoglossoides platessoides)

NW Atlantic

N/A

N/A

N/A

N/A

N/A

D

Yellow flounderl
(Limanda ferruginea)

NW Atlantic

20 971

24 273

18 948

21 447

20 803

F

Antarctic krill
(Euphausia superba)

Southern Ocean

114 426

104 182

125 987

117 728

118 165

U

Deepwater rose shrimp
(Parapenaeus longirostris)

Mediterranean

12 323

11 100

10 133

9 489

7 968

F

Blue and red shrimp
(Aristeus antennatus)

Mediterranean

1 904

2 209

2 011

2 063

2 083

N

Giant red shrimp
(family Aristeidae)

Mediterranean

4 463

1 833

1 768

2 409

1 546

N

Northern prawn
(Pandalus borealis)

NW Atlantic

232 028

230 757

253 691

282 692

354 907

F

West African geryon
(Chaceon maritae)

SE Atlantic

N/A

N/A

N/A

N/A

N/A

N

Deep-sea red crab
(Chaceon (Geyron) affinis)

NE Atlantic

N/A

N/A

N/A

N/A

N/A

N-O4

Patagonian squid
(Loligo gahi)

SW Atlantic

68 110

57 876

29 857

48 980

29 036

F

Squids
(family Loliginidae)

SE Atlantic

7 111

4 494

9 439

10 422

11 807

N

Neon flying squids
(Omastrephes bartrami)

SW Pacific

N/A

N/A

N/A

N/A

N/A

M

Argentine shortfin squid
(Illex argentinus)

SW Atlantic

940 054

750 203

540 357

503 624

129 279

F

Jumbo squid
(Dosidicus gigas)

SE Pacific

103 307

153 308

284 090

281 699

688 889

M

Jumbo flying squid
(Dosidicus gigas)

EC Pacific

103 307

153 308

284 090

281 699

688 889

M-F

Japanese flying squid
(Todarodes pacificus)

NW Pacific

570 427

528 523

504 438

487 576

447 363

M (R)

Sevenstar flying squid
(Martialia hyadesi)

SW Atlantic

653

115


37

59

N

Boreal clubhooksquid
(Onychoteuthis borealijaponica)

NW Pacific

N/A

N/A

N/A

N/A

N/A

F

Boreopacific armhook squid
(Gonatopsis borealis)

NW Pacific

N/A

N/A

N/A

N/A

N/A

F

Octopus
(family Octopodidae)

SE Atlantic

305

129

291

209

144

N

1The catch and state of exploitation is that of the species in the given area, with no distinction being made for straddling and not straddling (entirely EEZ) stocks, or for catches within EEZs or in the high seas

2Catch data from FAO FISHSTAT Plus

3Symbols: N = Not known; U = Underexploited; M = Moderately exploited; F = Fully exploited; O = Overexploited; D = Depleted; R = Recovering

4State of exploitation cannot be assessed with regard to standard criteria, considered non-sustainable by ICES (2005)


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