An annotated and illustrated catalogue of hake species known to date

FAO Species Catalogue for Fishery Purposes No. 2

HAKES OF THE WORLD
(Family Merlucciidae)

AN ANNOTATED AND ILLUSTRATED CATALOGUE OF HAKE SPECIES KNOWN TO DATE

by
D. Lloris
Instituto de Ciencias del Mar (CMIMA-CSIC)
Barcelona, Spain
J. Matallanas
Facultad de Ciencias
Universidad Autónoma de Barcelona
Bellaterra, Barcelona, Spain
and
P. Oliver
Instituto Español de Oceanografia
Palma de Mallorca, Spain


FOOD AND AGRICULTURE ORGANIZATION OF THE UNITED NATIONS

Rome, 2005

Table of Contents


The designations employed and the presentation of material in this publication do not imply the expression of any opinion whatsoever on the part of the Food and Agriculture Organization of the United Nations concerning the legal status of any country, territory, city or area or of its authorities, or concerning the delimitation of its frontiers or boundaries.

ISBN 92-5-104984-X

All rights reserved. Reproduction and dissemination of material in this information product for educational or other non-commercial purposes are authorized without any prior written permission from the copyright holders provided the source is fully acknowledged. Reproduction of material in this information product for resale or other commercial purposes is prohibited without written permission of the copyright holders. Applications for such permission should be addressed to the Chief, Publishing and Multimedia Service, Information Division, FAO, Viale delle Terme di Caracalla, 00100 Rome, Italy or by email to copyright@fao.org

© FAO 2005

Lloris, D.; Matallanas, J.; Oliver P.
Hakes of the world (Family Merlucciidae). An annotated and illustrated catalogue of hake species known to date.
FAO Species Catalogue for Fishery Purposes. No. 2. Rome, FAO. 2005. 57p. 12 colour plates.

ABSTRACT

This is a worldwide catalogue of the family Merlucciidae. Two subfamilies, Macruroninae and Merlucciinae, are recognized comprising four genera, Lyconodes, Lyconus, Macruronus and Merluccius, and 18 species. The following subspecies are proposed: Macruronus novaezelandiae magellanicus Lönnberg, 1907; Merluccius albidus magnoculus Ginsburg, 1954, Merluccius australis polylepis Ginsburg, 1954 and Merluccius polli cadenati Doutre, 1960 and Merluccius merluccius smiridus Rafinesque, 1810. The possibility of one other subspecies, Merluccius merluccius lessepsianus, represents the first record of Merluccius from the Red Sea. Merluccius paradoxus is first recorded from Madagascar.

In the introductory chapters, Merlucciidae systematics is debated, justifications for the proposed taxonomic organization are provided, and the characters used for the identifications are discussed.

Dichotomous keys are provided in the systematics chapter, enabling the identification of the hakes to the species level. Subfamilies and genera are also defined. The species are arranged in alphabetical order under each subfamily and genus to which they belong. The scientific name appears in bold at the head of each genus and species description, followed by the author, year of first description, and publication. Existing synonyms and FAO common names in English, French and Spanish are also provided. For each species there is an illustration followed by ten sections: description or diagnosis with differential characters; additional information; geographical distribution; habitat and biology; size; fisheries; fishery statistics; state of resources; local names, if any; and bibliographical references (author and year).

The review is completed by a series of colour plates showing details of different elements (heads, otoliths, hyomandibulars, urohyals) for a quick and efficient diagnosis of the genera and species of Lyconus, Macruronus and Merluccius.


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TABLE OF CONTENTS


FAO Species Catalogue for Fishery Purposes No. 2pdf

PREPARATION OF THIS DOCUMENT
ABSTRACT
ACKNOWLEDGEMENTS

1. INTRODUCTION pdf

1.1 Background
1.1.1 Merlucciidae Systematics
1.1.2 Character Analysis at the Genus Level
1.2 Characters Subject to Variability
1.3 Diagnostic Features of the Family

2. SYSTEMATIC CATALOGUEpdf

2.1 Key to Subfamilies, Genera and Species (except the genus Merluccius)
2.2 Subfamily: Macruroninae


Lyconodes
Lyconodes argenteus

Lyconus
Lyconus brachycolus
Lyconus pinnatus

Macruronus
Macruronus capensis
Macruronus novaezelandiae

2.3 Subfamily: Merlucciinae
2.4 Key to Species and Subspecies of the Genus Merluccius

Merluccius
Merluccius albidus
Merluccius angustimanus
Merluccius australis
Merluccius bilinearis

Merluccius capensis
Merluccius gayi
Merluccius hubbsi
Merluccius merluccius

Merluccius paradoxus
Merluccius patagonicus
Merluccius polli
Merluccius productus
Merluccius senegalensis

3. LIST OF SPECIES BY MAJOR FISHING AREAS pdf

4. BIBLIOGRAPHY pdf

5. INDEX OF SCIENTIFIC AND VERNACULAR NAMES pdf

6. COLOUR PLATESpdf