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R. K. Datta Central Sericultural Research & Training Institute,
Central Silk Board,
1. Area cultivated with mulberry in India: Mulberry foliage is the only food for the silkworm (Bombyx mori) and is grown under varied climatic conditions ranging from temperate to tropics. Mulberry leaf is a major economic component in sericulture since the quality and quantity of leaf produced per unit area has a direct bearing on cocoon harvest. In India, most states have taken up sericulture as an important agro-industry with excellent results. The total acreage of mulberry in India is around 282,244 ha. The details of area under mulberry cultivation in different states in India is shown in the table given below also in the Map: Table-1: Area under mulberry cultivation in different states
2. Species and varieties under cultivation in india There are about 68 species of the genus Morus, the majority of them occur in Asia, especially in China (24 species) and Japan (19). Continental America is also rich in its Morus species. The genus is poorly represented in Africa, Europe and Middle East, and it is not present in Australia. In India, there are many species of Morus, of which Morus alba, M. indica. M. serrata and M. laevigata grow wild in the Himalayas. Several varieties have been introduced belonging to M. multicaulis, M. nigra, M. sinensis and M. phillippinensis. Most of the Indian varieties of mulberry belong to M. indica. Table-2 : Mulberry varieties under cultivation in different states
In China there are 15 species. Out of which, 4 species, Morus alba, M. multicaulis, M. atropurpurea and M. mizuho are cultivated for sericulture. In the ex-USSR Morus multicaulis, M. alba, M. tartarica and M. nigra are present. Though mulberry cultivation is practiced in various climates, the major area is in tropical zone covering Karnataka, Andhra Pradesh and Tamil Nadu states, with about 90%. In the sub-tropical zone, West Bengal, Himachal Pradesh and north-eastern states have major areas under mulberry cultivation. The details of mulberry varieties under cultivation in different states of India is given in Table-2. 3. General Description Mulberry is a fast growing deciduous woody perennial plant. It has a deep-root system. The leaves are simple, alternate, stipulate, petiolate, entire or lobed. Number of lobes varies from 1 to 5. Plants are generally dioecious. Inflorescence is catkin with pendent or drooping peduncle bearing unisexual flowers. Inflorescence is always auxiliary. Male catkins are usually longer than the female catkins. Male flowers are loosely arranged and after shedding the pollen, the inflorescence dries and falls off. Number of parianth lobes are 4. Number of stamens are 4 and implexed in bud. Female inflorescence is usually short and the flowers are very compactly arranged. Number of parianth lobes are 4 and persistent. Ovary is one-celled and stigma is bifid. The chief pollinating agent in mulberry is wind. Fruit is a sorosis and the colour of the fruit is mainly violet black. Most of the species of the genus Morus and cultivated varieties are diploid having 28 chromosomes. However, triploids (2n=(3x)=42) are also extensively cultivated for their adaptability, vigorous growth and quality of leaves. 4. Climatic Requirement of Mulberry Mulberry thrives under various climatic conditions ranging from temperate
to tropic located north of equator between 28° N to 55°N latitude.
The ideal range of temperature is from 24-28°C. It grows well in places
with annual rainfall ranging from 600mm to 2500mm. In area with low rainfall,
the growth is limited due to moisture stress resulting in low yields. On
an average mulberry requires 85,000 gallons of water per hectare once every
10 days in case of loamy soils and 15 days in clayey soils. The atmospheric
humidity in the range of 65-80% is ideally suited for mulberry growth.
Sunshine is one of the important factors controlling growth and leaf quality.
In tropics, mulberry grows with a sunshine range of 9.0 to 13.00 hours
a day. Mulberry can be cultivated from sea level up to an elevation of
1000 m above mean sea level.
5. Soil Condition: Mulberry flourishes well in soils which are flat, deep, fertile, well drained, loamy to clayey, porous with good moisture holding capacity. The ideal range of soil pH is 6.2 to 6.8. The optimum pH required for mulberry is 6.5 to 6.8. Soil amendments may be used to correct the soil to get required pH. The quantity of gypsum or lime to be applied in different cases are as mentioned below: Soil amendments: (To be taken up after soil test)
pH range quantity of lime/ha (to bring pH to 6.8)
Method of application: The powdered gypsum/lime is mixed well
with soil with soil of the garden and irrigated to stagnation for 48-72h.
Later the water is leached out by drainage and dried (suitable for ploughing
and intercultural operations).
A) Mulberry Cultivation in South India under Rainfed Conditions (Karnataka, Andhra Pradesh and Tamil Nadu) I. Suitable Mulberry Varieties Kanva-2, S-13 and S-34 varieties are recommended for rainfed (rainfall
: 500-800 mm) regions of South India.
Kanva-2 : Belongs to Morus indica - Diploid. Widely cultivated in Southern India. Selection from natural population of Mysore Local variety. Flower and sorosis: Female, profuse flowering, many sorosis. Economic characters: Medium leaf maturity, yields about 30 to 35 MT/ha/year under irrigated condition. Leaf moisture content 70%, protein content 21% and sugar content 11.5%. High rooting ability (80%) and wide adaptability. Resistant to leaf spot. Moderately resistant to leaf rust and powdery
mildew.
S-13 : Belongs to M. indica. Selection from open pollinated hybrids of Kanva-2. Recommended for rainfed areas of South India during 1990. Flower: Male, profuse flowering. Economic characters: Yields 8-12 MT/ha/yr under rainfed conditions depending on rainfall. Moisture content 70.6% protein content 24.3% and sugar content 13.8%. Resistant to leaf spot and powdery mildew, moderately resistant to leaf
rust and tukra infestation.
S-34 : Belongs to M. india - Diploid. Selection from progeny of S30 x Berc 776. Recommended during 1990 for rainfed areas with black cotton soils of South India. Flower and sorosis: Male, profuse flowering, occasionally few sorosis. Economic characters: Under rainfed conditions, yields about 15 MT/ha/yr. Moisture content 70%, crude protein content 23.7%. Soluble sugar content 13.2%. Resistant to powdery mildew and leaf rust. Moderately resistant to leaf
spot and susceptible to tukra infestation.
II. Establishment of Mulberry i) Preparation of land: The land meant for mulberry cultivation is ploughed deep with heavy mould board plough up to a depth of 30-35 cm. Thereafter the land is repeatedly ploughed 2-3 times with a country plough to bring the soil to a fine tilth. The land should be properly levelled if it is sloppy. A basal dose of well decomposed Farm Yard Manure or compost is applied at the rate of 10 MT/ha and thoroughly incorporated into the soil. ii) Spacing: The spacing commonly followed for a rainfed garden is 90 x 90cm Pits of 35 x 35cm. are prepared. About 1 kg farm yard manure/pit should be added. iii) Preparation of cutting and planting: Branches of 8-10 months old and about 50 mm. in diameter should be used for preparation of cuttings of 22-25 cm. length with 5-6 healthy buds. Three cuttings are planted/pit in a triangular form with a spacing of 15 cm. leaving only one bud exposed above soil surface. If planting is done with saplings, then one sapling is sufficient/pit. Planting should be done during June /July after onset of monsoon. iv) Inter cultivation: During first year of plantation, inter-cultivation should be done manually. Once the mulberry plants are established, bullock ploughing is carried out. Application of fertilizer: 50N:25P:25K Kg/ha/yr in two doses.
v) Pruning and leaf harvest: First crop should be taken 6 months after plantation when mulberry gets well established. Then onwards, two more crops are taken during first year by leaf picking method. Mulberry should be pruned after the completion of one year at the onset of next monsoon. The pruning is carried out with a sharp sickle or a pruning saw at the height of 25-30 cm. from the ground. vi) Green manuring and cover mulching: Green manure crops can
be grown as inter-crop with mulberry. It should be done during monsoon
only. The green manure crops (cow pea, horse gram, dhaincha) should be
incorporated into soil by ploughing before the flowering starts and well
before rains cease. After that the plots may be given cover mulching with
any dry material or weeds in which seed is not a source of multiplication.
III. Maintenance of Mulberry Under Rainfed Condition (Second year onwards) Inputs required for rainfed garden (per ha. per year) Recommended inputs for gardens maintained under rainfed condition. Spacing : 90 cm. x 90 cm. 1. FYM/compost, 10 MT, single dose at the onset of monsoon
Crops like horse gram, cow pea, sun hemp, dhaincha, etc., should be incorporated into soil by ploughing before flowering and cessation of monsoon. LEAF HARVEST: Individual leaf harvesting should be made. The yield for different varieties is as mentioned below: K-2 – 10-12 MT/ha/yr
B) Mulberry Cultivation in South India under Irrigated Condition I. Suitable Mulberry Varieties Kanva-2, S-36, S-54, DD, MR2 (especially in Tamil Nadu) and V-1 varieties
are recommended for irrigated conditions.
Kanva-2 : Belongs to Morus indica - Diploid. Widely cultivated in Southern India after the same was recommended for cultivation in 1969 by CSR & TI, Mysore. Selection from natural population of Mysore Local variety. Flower and sorosis: Female, profuse flowering, many sorosis. Economic characters: Medium leaf maturity, yields about 30 to 35 MT/ha/year under irrigated condition. Leaf moisture content 70%, protein content 21% and sugar content 11.5%. Resistant to leaf spot. Moderately resistant to leaf rust and powdery
mildew. High rooting ability (80%) and wide adaptability.
S-36 : Belongs to M. indica. Developed at CSRTI, Mysore and recommended during 1984. Evolved from Berhampore local by chemical mutagenesis. Cultivated in Southern India. Moderate rooting ability. Economic characters: Yields about 38 - 45 MT/ha/yr under assured irrigated conditions of South India. Moisture content 76 % protein content 22 % and carbohydrate content 28 %. Tolerant to leaf spot and powdery mildew. Moderately susceptible to leaf rust and to tukra infestation. Most suitable to young age silkworm rearing. Sapling plantation recommended
due to moderate rooting ability.
S-54 : Belongs to M. india. Developed at CSRTI, Mysore and recommended during 1984. Selected from Berhampore local by chemical mutagenesis (EMS). Recommended for assured irrigated conditions of South India. Economic characters: Yields about 45 MT/ha/yr under assured irrigated
conditions. Highly responsive to agronomical input. Moisture content of
leaf 70.5% protein content 23.9% and sugar content 13.8%. Moderately resistant
to powdery mildew and leaf rust and resistant to leaf spot. Looses moisture
very quickly. Good rooting ability.
DD : Selected from natural population of Dehra Dun variety and recommended by Karnataka State Sericultural Research and Development Institute, Thalaghattapura. Recommended for Southern India. Morphology: Erect, thin branches, coarse leaves, greenish grey colour
bark. Lower branches spreading, leaves unlobed, big size, ovate shape.
Yields 35-40 MT/ha/yr under assured irrigation condition.
MR – 2 : Belongs to M. sinensis – Diploid. Selection from open pollinated hybrid population. Developed at CSR & TI, Mysore and recommended for propagation in Tamil Nadu. Mainly cultivated in Tamil Nadu under both irrigated condition in plains and rainfed conditions in hilly regions. Economic characters: Yields about 30-35 MT/ha/yr under irrigated conditions of Tamil Nadu. Moisture content 68%, protein content 23.2%, sugar content 13.2% (Fig. 7). Resistant to powdery mildew disease, suitable for hilly areas.
V-1 : Belongs to M. indica L. Recently developed from a cross of S-30 and Berc.776 at CSR&TI, Mysore. Recommended during 1996 for assured irrigated conditions. Flower: Male, profuse flowering, occasionally few sorosis. Economic characters: Yields about 70 MT/ha/yr under assured irrigated conditions. Very high sprouting. Moisture content 78.9% and 72.5% in young and matured leaves respectively, protein content 24.6% and total sugar content 16.98%. Moderately resistant to leaf rust and tukra infestation and resistant
to leaf spot. Quick sprouting ability and very high rooting ability (>
94%) high photosynthetic rate and higher water use efficiency are additional
advantages. Moreover, leaves are suitable for both young and grown bivoltine
silkworm rearing.
II. Establishment of Mulberry i) Selection of site: Mulberry flourishes well in soils which are flat, deep, fertile, well drained, loamy to clayey, porous with good moisture holding capacity. The ideal range of soil pH is 6.2 to 6.8. Mulberry can be grown in saline, alkaline and acidic soils after suitably amending the soils. ii) Preparation of land: The land meant for mulberry cultivation is ploughed deep with heavy mould board plough up to a depth of 30 – 35 cm. Thereafter the land is repeatedly ploughed 2-3 times with a country plough to bring the soil to a fine tilth. The land should be properly leveled if it is sloppy. A basal dose of well decomposed farm yard manure or compost is applied at the rate of 20 MT/ha and thoroughly incorporated into the soil. iii) Plantation can be raised by using both cuttings or saplings. The varieties ideally suited for irrigated conditions are Kanva-2, S-36 and V-1. Branches of 6-9 months old and about 15 ml.. in diameter should be used for preparation of cuttings of 15-18 cm. length having 3-4 healthy buds for raising nursery or for planting directly in the field. iv) Spacing: Plant spacing of 90 x 90 cm. is ideal for mulberry. While taking Plantation directly, two cuttings/pit required to be used. In case of using saplings, only one sapling/pit is sufficient. Presently a paired row plantation with the spacing of (90+150)cm x 60cm, is recommended. v) Inter-cultivation: Two months after planting, weeding is done. A second weeding is done after another 2-3 months. Thereafter, inter-cultivation should be done after every shoot or leaf harvest. vi) Irrigation: The plantation should be taken up during the onset of monsoon to take advantage of the rains. If the rain is not sufficient, the land should be irrigated at regular intervals of 8-14 days depending on type of soil. About one and a half to two acre inches of water is required/irrigation. vii) Application of fertilizer: The total dose of fertilizer to be applied in first year is 100 N: 50 P: 50 K/kg/ha/yr. This is applied in two doses. The first dose is applied when the plantation is about 2 months old at the rate of 50 N: 50 P: 50 K/kg/ha. The second dose is applied after taking the leaf harvest at the rate of 50 kg N alone/ha. viii) Pruning: After 6 months of plantation, mulberry attains
a height of about 1.50 to 1.75 m. and is ready for harvest. The first harvest
is taken by bottom pruning. The second leaf harvest is taken after 12 weeks
of first leaf harvest and the third harvest is taken 12 weeks after the
second harvest by shoot harvest. From second year onwards, harvest is made
at an interval of 70 days by shoot harvest method.
III. Maintenance of Mulberry under Irrigated Conditions (Second year onwards) Spacing 90 cm. x 90 cm. or (90 + 150) cm x 60 cm Recommended inputs:
** VAM inoculation is not required for the garden, planted with Mycorrhiza inoculated saplings. *** 1st spray: In between 10-15 days after pruning/leaf plucking 2nd spray : 10 days after Ist spray.
C. Mulberry Cultivation in Hilly Areas I. Suitable Mulberry Varieties S-1, S-7999, S-1635, S-146, Tr-10 and BC-259 varieties are recommended
for hilly regions of north and north-eastern India.
II. Establishment of Mulberry i. Preparation of land: If the land is having gentle slope, it can be leveled with minor land shaping and providing suitable type of bunds across the slope. If the slope is more, contour bunding terrace planting or contour line planting can be adopted. In more sloppy areas, platform for individual plants on contour lines is more suitable as the same involves less soil cutting. ii. Spacing: Spacing for tree planting depends on soil topography, extent of land available for cultivation and training method. In case of gentle slope, 3`X 3`, 5’x 5’ may be adopted. In case of more sloppy land 10’ x 10’ can be adopted. Pits are to be prepared for plantation. In case of deep textured loose soils, 45 x 45cm. and in had shallow soils 60cm x 60cm x 60cm) pits are to be prepared. For each pit, 5kg (one iron pan) of FYM or compost must be applied. iii. Planting: Saplings of 5 months age with 5-6 feet is found suitable to plant during the regular onset of monsoon. One sapling/ pit should be planted. The saplings should be supported with stick to ensure straight growth. iv. After care of plantation: After one month, all the buds except
top 5-6 should be removed carefully without damaging the bark. Weeds around
the plant should be removed and regular pot watering should be given. After
3 months of planting, second weeding should be given and 25g of suphala/plant
should be applied in trench and should be covered with soil. 2nd
dose of fertilizer (25g urea/plant) should be given before cessation of
monsoon. Plants must be protected from grazing.
III. Recommendation with Institute Technologies for Hilly Areas (Tropical high land) Spacing 90 cm. x 90 cm.
CAN: Calcium Ammonium Nitrate
Since the recommendation is general, the inputs like fertilizers and
amendments may be applied on the basis of soil test reports.
D. Inputs for Mulberry Cultivation in Temperate and Sub-Temperate Regions The following varieties are found suitable for temperate and sub-temperate regions. Sub-temperate - Chak majra, S-146
The leaf yield of different varieties are as mentioned below: China white 15-20 MT/ha/yr Chak majra 20-22 MT/ha/yr Mulberry can be successfully grown as an inter-crop (medium mixed tree) between the rows of tea/coffee as shade plants. By this besides providing shade to the tea/coffee plants, good quantity of quality leaves can be obtained for silkworm rearing and as feed for cattle and goats. Further, the pruned shoots are a good source of firewood. Mulberry can also be grown as an inter-crop for cultivation in the wider
space existing between sources of coconut plantation. A survey has shown
that mulberry is inter-cropped with coconut in Channapatna, Ramanagaram,
Kanakapura and Bangalore areas of Karnataka state.
7. Insecticide/ Pesticide/
Fungicide and Other Inputs (ha/y) (if required)
8. Animal Feeding Practices and Other Traditional Uses Mulberry is appropriately known as "Kalpa Vruksha" as all the parts of the plant has many uses. It is essential to sericulture as the foliage constitute the sole feed of mulberry silkworm Bombyx mori. Mulberry (Morus spp) is a fast growing tree which for convenience of sericulture practices is maintained as a bush. It produces very large amounts of renewable bio-mass in the form of branches, shoots, leaves and fruits. If mulberry is used for silkworm rearing it is possible to get 30-35 MT/ha of leaf every year. By growing mulberry, a farmer gets fodder, fuel and fertilizer. With regard to fodder for animals, farmers in India feed their cows and goats with left over branches and leaves during silkworm rearing period. Many farmers feed their animals with full twig of mulberry when leaves are in excess. But, they always mix mulberry twigs with straw. Farmers also use the mulberry twigs as their fuel source after pruning of the garden. Left over twigs are allowed to dry in the garden itself. Refuses of rearing are also converted to valuable Farm Yard Manure for mulberry garden by the farmers by putting it in a pit and keeping it for 4-5 months prior to its use. As mulberry is mainly propagated by cuttings in tropics and sub-tropics, certain quantity of pruned branches can be used for preparation of cuttings and the remaining as fuel wood. One hectare of mulberry garden yields about 12.1 MT of mulberry sticks. The energy generated/ha (50% moisture loss) is 27830 Kcal (@ 4600 calories/kg of mulberry wood). Accordingly, mulberry can be exploited by raising it as "energy plantation" in cultivable/wasteland/low lying areas/canal bund/road side/fringe areas of the forest etc., under various afforestation, watershed development and soil conservation programmes. Traditional uses of mulberry: The various species of the genus Morus viz., M. laevigata, M. serrata, M. alba and M.indica has varied uses which are enumerated below: M. laevigata: The trees of this species produce sweet fruits which are used in juice and jam making in central India. In North-east India the wood is utilized as firewood, house building, furniture making, for making stocks, spokes, poles, shafts of carriages and casts. The wood is suitable for plywood making and paneling, carving and making of toys and tea chests. It is used for making tennis rackets. The straight log of the tree is used as support in house building work. M. serrata: The wood is used for furniture making and carving, toys making, sports goods, agricultural implements and cheap types of rifles and guns. M. alba: It is cultivated in the hilly and plain areas of India (Himalayan region) for silkworm rearing. It is also used as a avenue tree and in social forestry. Fruits are made into juice, liquor and stews. The wood finds use in sports goods industry. It is also used for house building, agricultural implements, furniture, for making spokes, poles, shafts and bent parts of carriages and carts. The stem bark is used for paper making. M. indica: The cultivated forms belong to M. indica which are utilized in silkworm rearing. There are few profuse fruiting varieties occurring in Maharastra and Meghalaya which can be utilized as female parent in breeding programmes. The fruits are used for jam, jelly and juice making in Maharashtra. The pruned branches are used as fuel. Medicinal uses: The various parts of mulberry plant finds use
in ayurvedic preparations. The leaves have diaphoretic and emollient effects.
The leaves are used for making a decoction which can be used as a gargle
to get relief from throat inflammation. The fruits are used to treat sore
throat, depression, high fever and is both a coolant and laxative. The
roots extract have hypoglycemic properties. The root bark is used as an
anthelmintic, purgative and vermifuge. Mulberry root juice is administered
to patients with high blood pressure. The chinese use the leaf tips from
young leaves to boil with tea and use it to control blood pressure. The
milky latex is used as a plaster for sores and for preparation of dermal
creams.
9. Fruiting Habit In South India, fruits are observed in two seasons in a year once during
October – November and the other during March – May. However, whenever
mulberry is pruned or defoliate flowering takes place along with sprouting
of auxiliary buds followed by fruit formation. This feature of mulberry
is utilized in mulberry breeding programmes. The immature fruits are green
in colour but changes to purplish to violet black. In certain species like
Morus
alba the fruits are white to pinkish and very sweet. In Morus lavigata
the fruits are very long sometime measuring up to 18cm.
10. Leaf Chemical Composition Chemical composition of leaf varies with variety and maturity. However
on the basis of the analysis carried out at CSR& TI Mysore , the chemical
composition of the leaf is as follows:
11. Main Pests and Diseases of Mulberry Key pests of mulberry: Diaphania pulverulentalis – Leaf roller. Spilarctia obliqua – Bihar hairy caterpillar (Sporadic pest).
Thrips, Jassids, Scale insects, short horned grasshopper. Tolerance of varieties to pests: The tukra incidence in rainfed areas was found to be maximum in S-34
variety followed by MR-2, BERC 776, MS7 and S-13 (Srinivas et. al., 1996).
The spread of tukra in V-1 variety is less compared to other varieties
viz., Local, K-2, S-13, S-34, S-36 suggesting V-1 variety to be relatively
tolerant to tukra (Anonymous, 1998, Sujatha, 1997). Screening of germplasm
maintained at CSRTI, Mysore, indicated the variety TOGHWASE – Acc.No. 257
was found to be tolerant to pest attack (unpublished data).
Main diseases of mulberry:
12. References Choudhury, P.C., 1997. Mulberry cultivation, CSRTI, Mysore. Cultivation technology for chawki mulberry garden. Agronomy Department, CSRTI, Mysore. Chemical composition and quality of mulberry leaves. Text book of tropical sericulture, 1976 pp:154 - 165. Handbook of silkworm rearing, 1972 – Agriculture Technique Manual 1, Fuji publishing co., ltd., Tokyo, Japan. Jalaja S.Kumar, Sarkar, A., Datta, R.K. A breakthrough in mulberry breeding in sustainable cocoon production. In global silk scenario - 2001, Proc. Of the International Conference on Sericulture. Oxford and IBH publishing Co. Pvt.Ltd., pp. 242-247. Jolly, M.S., 1987. Appropriate sericulture techniques, ICTRETS, Mysore. Rangaswami, G. Narasimhanna, M.N., Kasi Viswanathan K. and Sastry, C.R., 1976. Manual on Sericulture Vol.1, FAO, Rome. Ravindran, S., Ananda Rao, A., Girish Naik, V., Tikader, A., Mukherjee, P. and Thangavelu, K. 1997. Distribution and variation in mulberry germplasm. Indian J. plant genet. Resources 10 (2):233-242. Ravindran, S., Tikader, A., Girish Naik, V., Ananda Rao, A. and Mukherjee, P., 1988. Distribution of mulberry species in India and its utilization. Poster paper presented in "National Dialogue". Sastry, C.R., 1984. Mulberry varieties, exploitation and pathology, Sericologia, 24 (3): p.333-359. Sarkar, A. and Jalaja S.Kumar, 1996. Gradual improvement of mulberry varieties under irrigated conditions in South India and potential programme for varietal selection. Internationa Symposium on sericulture science, Hangzhhou, China. Sarkar, A., Jalaja S.Kumar and Datta, R.K., 1999. Potentiality of Victory-1 under irrigated conditions of South India. Indian Silk, May, 1999. Sarkar, A., Jalaja S.Kumar and Datta, R.K. Gradual improvement of mulberry varieties under irrigated condition in South India and the optimal programme for varietal selection in the tropics. Sericologia (In Press). Sericulture: New technologies, 1996, CSRTI, Mysore. Susheelamma, B.N., Jolly, M.S., Sengupta, K., Giridhar, K., Baksh, S., Mogili, T., Mallikarjuna, R.S. and Pavan Kumar, T., 1992. Statistical analysis of adaptability of drought resistant mulberry genotypes. Sericologia 32 (4) : 619-628. Ullal, S.R. and Narasimhanna, M.N., 1987. Handbook of Practical Sericulture, Central Silk Board, Bangalore. Watt, G. 1981. Dictionary of economic products of India Vol.5, pp.279-284, Periodical Experts, Delhi. |
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