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PART 1 - INTRODUCTION AND TABLES

Introduction

This paper gives the values of the yield of edible tissue and the protein and fat contents of the world's commercially more important fish and shellfish species, derived from a survey of published, and some unpublished, material. The information is intended to assist FAO in estimating the contribution of fish to the nutrition of individual countries from their data on catches, landings and disposal. Much of the data collected is insufficiently comprehensive, especially in relation to yield, to allow definitive estimates to be made: many of the selected values are considered provisional, or tentative.

Data of the yield and composition (as a percentage of the edible portion) of the species surveyed are presented in Table 1. Much of the published data on which Table 1 is based is of less value than it could be, because of the universal failure of authors to appreciate the many factors that can affect yield and composition. A high proportion of the individual measurements in the literature are from a small number a fish, sometimes only one, caught on a single occasion. The possible effects on both yield and composition of the date of catching, the location, the size of the fish and natural variability have selfom, if ever, been fully taken into account. Yield measurements are particularly variable: it is not impossible to find published yields for a species that vary by a factor of 2. Apart from the factors already mentioned as causing variability, all authors have failed to define exactly terms such as “fillet” and “edible portion”.

The separate yields of skinless fillets and total edible flesh allow some flexibility in assessing the actual amount of fish tissue consumed in particular countries. In many developed countries a high proportion of fish is eaten in the filleted condition, with no bones or with pinbones only present, and often without skin. The remaining edible tissue is consigned with the bones, offal and possibly skin to fish meal manufacture, when it may contribute only indirectly to human nutrition. The extent to which skin is eaten is not addressed here since reliable information is lacking. In other countries, including most developing countries, fish is sold to the consumer either in the whole, gutted condition, or in some dressed or split form. In these circumstances nearly all the muscle tissue will be eaten and the “edible flesh” column in Table 1 is an estimate of such consumption. In any particular case a decision will need to be made on which yield figure is most suitable. (An intermediate figure could be appropriate.)

Yields are given to the nearest whole number, protein and fat contents to the first decimal place and energy values to the nearest whole number.

Energy values have been calculated using the factors:

Protein4.27 kcal/g
Fat9.02 kcal/g
Glycogen4.11 kcal/g

1 kcal = 4.185 kJ

General notes on Table 1:

  1. Because of the inadequacy of the collected data many of the selected values of yield and some values of composition are considered tentative or provisional: these figures are in square brackets.

  2. For each listed species, the Monograph giving fuller details will be found in Part 3, either under the English name of the species or as indicated in a numbered footnote.

  3. The species or groups listed include, with a few exceptions such as species used mostly for reduction to fish meal, almost all those with an annual world catch exceeding 100,000 metric tons (MT) per year (together with some additional species that were considered at the same time.)

  4. Selected values of yields are given as percentages of the whole weight of the fish.

  5. Selected values of proteins and fat content are given as percentages of the edible tissue, muscle in the case of finfish, but including other organs in many shellfish.

  6. Where two or more figures are given in a single column, this is explained in a numbered footnote.

  7. The separate yields of skinless fillets and total edible flesh allow some flexibility in assessing the actual amount of fish tissue consumed in particular countries. In many developed countries a high proportion of fish is eaten in the filleted condition, with no bones or with pinbones only present, and often without skin. The remaining edible tissue is consigned with the bones, offal and possibly skin to fish meal manufacture, when it may contribute only indirectly to human nutrition. The extent to which skin is eaten is not addressed here since reliable information is lacking. In other countries, including most developing countries, fish is sold to the consumer either in the whole, gutted condition, or in some dressed or split form. In these circumstances nearly all the muscle tissue will be eaten and the ‘edible flesh’ column in Table 1 is an estimate of such consumption. In any particular case a decision will need to be made on which yield figure is most suitable. (An intermediate figure could be appropriate.)

  8. Yields are given to the nearest whole number, protein and fat contents to the first decimal place and energy values to the nearest whole number.

  9. Energy values have been calculated using the factors:

Protein4.27 kcal/g
Fat9.02 kcal/g
Glycogen4.11 kcal/g

1 kcal = 4.184 kJ

Comment on the exercise:

Much of the published data on which Table 1 is based, is of less value than it could be, because of the universal failure of authors to appreciate the many factors that can affect yield and consumption. A high portion of the individual measurements in the literature are from a small number of fish, sometimes only one, caught on a single occasion. The possible effects on both yield and composition of the date of catching, the location, the size of the fish and natural variability have seldom, if ever, been fully taken into account. Yield measurements are particularly variable: it is not impossible to find published yields for a species that vary by a factor of 2. Apart from the factors already mentioned as causing variability, all authors have failed to define exactly terms such as ‘fillet’ and ‘edible portion’.

As far as possible, the data have been obtained from original sources, not from other compilations or from abstracts.

Table 1

Yield and composition of surveyed species, in order of English name
SpeciesYield (as % of whole fish or shellfish weight)Composition (as % of edible portion)Energy Value
(see notes page 10)Skinless FilletEdible FleshMeatsProteinFatGlycogenKcalKjoules
Akiami paste shrimp  [100][16.2][1.3] [81][338]
Alaska pollack3641 16.60.6 76319
Albacore (1)[30]58 23.74.6 143597
Amberjacks nei[46][53] 22.02.3 115480
American cupped oyster(2)  108.51.82.764266
American plaice (3)3449 17.61.6 90375
Anchoveta (4,28)[51]62 18.0(20.2)2.8(4.2) 102(124)427(519)
Anchovies nei (4,28)[51]62 18.0(20.2)2.8(4.2) 102(124)427(519)
Argentine hake (5)4153 17.82.5 99412
Argentine shortfin squid (6)  6717.91.3 88369
Atka mackerel[34][54] [17.5][4.0] [111][464]
Atlantic cod3447 18.10.8 56234
Atlantic herring (29)4661 17.813.8(14.2/6.1) 200(204/131)839(854/548)
Atlantic horse mackerel (7)[46]52 19.75.8 136571
Atlantic mackerel5461 18.711.4 183764
Atlantic redfishes2945 18.42.7 103431
Bali sardinella (8)[53]65 20.24.5 127531
Banana prawn (9)  5720.51.3 99415
Bigeye tuna (1)[30]58 23.74.6 143597
Black Sea sprat (10)[50][56] [17.1][8.1] [146][611]
Blue mussel  2411.21.82.574311
Blue whiting2849 18.40.3 81340
Bombay duck-[57] [8.8][0.3] [40][169]
Calico scallop (11,30)  [15],[6][16.0],15.8[0.8],0.6[2.9],2.4[87],83[366],346
California pilchard (12)5053 18.97.9 152636
Cape hakes (5)4153 17.82.5 99412
Cape horse mackerel (7)[46]52 19.75.8 136571
Capelin[57][60] 14.95.9 117489
Central Pacific anchoveta (4,28)[51]62 18.0(20.2)2.8(4.2) 102(124)427(519)
Characins[38][57] [17.6][10.5] [170][711]
Chilean hake (5)4153 17.82.5 99412
Chilean jack mackerel (7)[46]52 19.75.8 136571
Chub mackerel4657 20.49.5 173723
Chum salmon (13)[50]62 20.56.3 144604
Chichlids nei (14)[37]37 17.62.1 94394
Clams nei (15)[18] 10.81.1 3.972302
Coho salmon (13)[50]62 20.56.3 144604
Common carp36[54] 17.54.7 117490
Common scallop (11,30)  [15],[6][16.0],15.8[0.8],0.6[2.9],2.4[87],83[366],346
Common squids (6)  6717.91.3 88369
Cunene horse mackerel (7)[46]52 19.75.8 136571
Cupped oysters nei (2)  108.51.82.764266
Cuttlefishes, bobtail squids (6)  [63]17.91.3 88369
Cyprinids nei (16)36[54]17.54.7  117490
Dagaas-[63] [16.2][3.1] [97][406]
Dentex seabreams, etc nei4054 19.92.2 105439
European anchovy (4,28)[51]62 18.0(20.2)2.8(4.2) 102(124)427(519)
European hake (5)4153 17.82.5 99412
European pilchard[50]62 20.08.1 158663
European plaice3449 17.61.6 90375
European sprat[50][56] [17.1][8.1] [146][611]
Filefishes (17)[33][36] 18.80.7 87362
Flathead mullet (18)3450 19.84.0 121505
Freshwater bream (16)36[54] 17.54.7 117490
Frigate and bullet tunas (1)[30]58 23.74.6 143597
Goldstripe sardinella (8)[53]65 20.24.5 127531
Green mussel (19)  2411.21.82.574311
Greenland halibut (3)3449 13.013.7 179749
Haddock3548 18.60.2 81340
Hairtails, cutlassfishes4659 20.13.9 121506
Hard clam (15)  [18]10.81.13.972302
Indian mackerel (20)[45]57 20.43.2 116485
Indian mackerels nei (20)[45]57 20.43.2 116485
Indian oil sardine (8)[53]65 20.24.5 127531
Indian scad (21)[52][52] 21.32.5 114475
Jack and horse mackerels nei (7)[46]52 19.75.8 136571
Jacks, crevalles nei[33]56 [19.8][1.7] [100][418]
Japanese amberjack (22,31)[46][53] 22.02.3(12.3) 115(205)480(857)
Japanese anchovy (4,28)[51]62 18.0(20.2)2.8(4.2) 102(124)427(519)
Japanese Manila) clam ((15)  [18]10.81.13.972302
Japanese claim (15)  [18]10.81.13.972302
Japanese flying squid (6)  6717.91.3 88369
Japanese jack mackerel (7)[46]52 19.75.8 136571
Japanese pilchard5053 18.97.91 52636
Japanese scad (21)[52][52] 21.32.5 114475
Japanese scallop (11,30)  [15],[6][16.0],15.8[0.8],0.6[2.9],2.4[87],83[366],346
Japanese Spanish mackerel (23)5767 20.0[4.4] [125][523]
Jellyfishes-- [0.8][0.1] [4]18]
Kawakawa (1)[30]58 23.74.6 143597
Largehead hairtail (24)4659 20.13.9 121506
Mediterranean horse mackerel (7)[46]52 19.75.8 136571
Milkfish[46]61 21.14.7 132554
Mullets nei3450 19.84.0 121505
Narrow-barred spanish        
king) mackerel ((23)5767 20.0[4.4] [125][523]
Nile perch[57][65] [21.2][0.8] [98][409]
North Pacific anchovy (4,28)[51]62 18.0(20.2)2.8(4.2) 102(124)427(519)
North Pacific hake (5)4153 17.82.5 99412
Northern bluefin tuna (1)[30]58 23.74.6 143597
Northern brown shrimp (9)  5720.51.3 99415
Northern prawn  [36]16.81.0 81338
Northern white strimp (9)  5720.51.3 99415
Ocean quahog (15)  [18]10.81.13.972302
Octopuses (6)  [79]17.91.3 88369
Pacific cod3041 17.30.7 80335
Pacific cupped oyster  108.51.82.764266
Pacific herring[52]60 17.811.6 181756
Pacific saury[52]64 21.88.6 171714
Penaeus shrimps nei  5720.51.3 99415
Pink salmon (13)[50]62 20.56.3 144604
Ponyfishes (Slipmouths) nei-[36] 17.9[1.0] [85][358]
Round sardinella (8)[53]65 20.24.5 127531
Saithe3447 18.30.5 83346
Sardinellas nei (8)[53]65 20.24.5 127531
Scads[52][52] 21.32.5 114475
Scallops nei (30)  [15],[6][16.0],15.8[0.8],0.6[2.9],2.4[87],83[366],346
Scorpionfishes, etc nei (25)2945 18.42.7 103431
Sea mussels nei (19)  2411.21.82.574311
Sea scallop (11,30)  [15],[6][16.0],15.8[0.8],0.6[2.9],2.4[87],83[366],346
Seerfishes nei5767 20.0[4.4] [125][523]
Short neck clams (15)  [18]10.81.13.972302
Silver hake (5)4153 17.82.5 99412
Silver seabream (26)4054 19.92.2 1.5439
Skipjack tuna[46]62 24.23.8 138576
Snoek (24)4659 20.13.9 121506
Sockeye salmon (13)[50]62 20.56.3 144604
South American pilchard (12)5053 18.97.9 152636
Southern African anchovy (4,28)[51]62 18.0(20.2)2.8(4.2) 102(124)427(519)
Southern African pilchard (12)5053 18.97.9 152636
Southern blue whiting (27)2849 18.40.3 81340
Southern bluefin tuna (1)[30]58 23.74.6 143597
Squids nei (6)  6717.91.3 88369
Stolephorus anchovies (4,28)[51]62 18.0(20.2)2.8(4.2) 102(124)427(519)
Striped venus (15)  [18]10.81.13.972302
Surf clam (15)  [18]10.81.13.972302
Threadsail filefish (17)[33][36] 18.80.7 87362
Tilapias nei[37]37 17.62.1 94394
Toli shad[50][60] 19.8[12.6] [198][829]
Triggerfishes, durgons[33][36] 18.80.7 87362
Whiting32[49] 18.2[0.4] [81][340]
Yellowfin sole (3)3449 17.61.6 90375
Yellowfin tuna (1)[30]58 23.74.6 143597

nei = not elsewhere included


Notes to Table 1

1MonographentitledTUNAS
2""PACIFIC CUPPED OYSTER
3""EUROPEAN PLAICE
4""ANCHOVIES
5""HAKES
6""SQUIDS, ETC
7""JACK AND HORSE MACKERELS
8""SARDINELLAS
9""PENAEUS SHRIMPS NEI
10""EUROPEAN SPRAT
11""SCALLOPS NEI
12""JAPANESE PILCHARD
13""PACIFIC SALMON
14""TILAPIAS NEI
15""CLAMS
16""COMMON CARP
17""TRIGGERFISHES, DURGONS
18""MULLETS NEI
19""BLUE MUSSEL
20""INDIAN MACKERELS
21""SCADS
22""AMBERJACKS NEI
23""SEERFISHES NEI
24""HAIRTAILS, CUTLASSFISHES
25""ATLANTIC REDFISHES
26""DENTEX, SEABREAMS, ETC, NEI
27""BLUE WHITING
28First figure in composition and energy columns is for whole fish, figure in brackets for flesh
29First figure in fat and energy columns is the weighted mean of the second and third figures; the second figure is for Atlantic-caught and third figure for Baltic-caught fish
30First figure refers to meat including roe, second figure to meat excluding roe
31First figure in fat and energy column is for wild, second figure for farmed fish


Table 2

Species items included in Survey, in order of taxonomic codes
Taxonomic Code Scientific Name English Name 1985 catch (metric tons)
1,21(05)xxx,xxStolothrissa, Limnothrissa sppDagaas116,397
1,21(05)001,05Clupea harengusAtlantic herring1,519,202
1,21(05)001,07Clupea pallasiPacific herring292,723
1,21(05)012,xxSardinella sppSardinellas nei847,767
1,21(05)012,03Sardinella gibbosaGoldstripe sardinella108,543
1,21(05)012,04Sardinella longicepsIndian oil sardine290,229
1,21(05)012,10Sardinella aurita (= anchovia)Round sardinella162,169
1,21(05)012,23Sardinella lemuruBali sardinella54,058
1,21(05)013,01Sardinops melanosticta (-us)Japanese pilchard4,722,862
1,21(05)013,02Sardinops caerulea (-us)Californian pilchard372,344
1,21(05)013,03Sardinops sagaxSouth American pilchard5,814,448
1,21(05)013,05Sardinops ocellata (-us)Southern African pilchard89,798
1,21(05)034,04Hilsa toli (= Tenualosa toli)Toli shad146,754
1,21(05)059,01Clupeonella delicatula (= cultriventris)Black Sea sprat (Clupeonella)402,889
1,21(05)064,01Sardina pilchardusEuropean pilchard921,061
1,21(05)066,01Sprattus sprattusEuropean sprat266,829
1,21(06)xxx,xxEngraulidaeAnchovies183,902
1,21(06)002,01Engraulis encrasicolusEuropean anchovy598,798
1,21(06)002,02Engraulis japonicusJapanese anchovy349,336
1,21(06)002,07Engraulis mordaxNorth Pacific anchovy153,723
1,21(06)002,08Engraulis ringensAnchoveta (Peruvian anchovy)986,796
1,21(06)002,12Engraulis capensisSouthern African anchovy323,239
1,21(06)015,03Cetengraulis mysticetusCentral Pacific anchoveta245,264
1,21(06)050,xxStolephorus sppStolephorus anchovies244,181
1,22(02)001,01Chanos chanosMilkfish336,518
1,23(01)009,02Oncorhynchus gorbuschaPink (Humback) salmon301,086
1,23(01)009,03Oncorhynchus ketaChum (Keta, Dog) salmon267,612
1,23(01) 009,06Oncorhynchus nerkaSockeye (Red) salmon150,794
1,23(01) 009,08Oncorhynchus kisutchCoho (Silver) salmon48,185
1,23(04) 002,01Mallotus villosusCapelin2,215,527
1,32(11) 001,02Harpadon nehereusBombay duck119,229
1,38(01) xxx,xxCharacidaeCharacins154,452
1,40(02) xxx,xxCyprinidaeCyprinids nei186,018
1,40(02) 001,02Abramis bramaFreshwater bream58,020
1,40(02) 002,01Cyprinus carpiocommon carp490,438
1,47(02) 001,01Cololabis sairaPacific saury273,760
1,48(04) 002,02Gadus morhuaAtlantic cod1,947,291
1,48(04) 002,11Gadus macrocephalusPacific cod409,598
1,48(04) 010,01Melanogrammus aeglefinusHaddock377,548
1,48(04) 015,01Pollachius virensSaithe (pollock)504,948
1,48(04) 016,01Theragra chalcogrammaAlaska pollak6,132,334
1,48(04) 033,01Micromesistius poutassouBlue whiting660,619
1,48(04) 033,02Micromesistius australisSouthern blue whiting106,199
1,48(04) 034,01Merlangius merlanguswhiting157,049
1,48(05) 004,xxMerluccius capensis, paradoxusCape hakes541,815
1,48(05) 004,01Merluccius merlucciusEuropean hake131,774
1,48(05) 004,04Merluccius bilinearisSilver hake99,135
1,48(05) 004,05Merluccius gayiChilean hake47,064
1,48(05) 004,06Merluccius hubbsiArgentine hake371,690
1,48(05) 004,07Merluccius productusNorth Pacific hake99,297
1,65(01) xxx,xxMugilidaeMullets nei162,348
1,65(01) 001,02Mugil cephalusFlathead mullet47,734
1,70(01)167,07Lates niloticusNile perch152,568
1,70(23)004,xxTrachurus sppJack and horse mackerels nei390,374
1,70(23)004,01Trachurus trachurusAtlantic horse mackerel209,440
1,70(23)004,03Trachurus japonicusJapanese jack mackerel173,775
1,70(23)004,05Trachurus murphyiChilean jack mackerel2,148,841
1,70(23)004,08Trachurus mediterraneusMediterranean horse mackerel120,319
1,70(23)004,13Trachurus capensisCape horse mackerel455,421
1,70(23)004,14Trachurus trecaeCunene horse mackerel56,766
1,70(23)043,xxDecapterus sppScads318,669
1,70(23)043,07Decapterus maruadsiJapanese scad71,666
1,70(23)043,08Decapterus russelliIndian (Russell's) scad55,201
1,70(23)044,xxCaranx sppJacks, crevalles, nei102,318
1,70(23)048,xxSeriola sppAmberjacks nei42,035
1,70(23)048,02Seriola quinqueradiataJapanese amberjack150,961
1,70(35)xxx,xxLeiognathidaePonyfishes (slipmouths)nei159,471
1,70(39)xxx,xxSparidaeDentex (Porgies), seabreams, etc nei102,866
1,70(39)121,08Chrysophrys majorsilver seabream44,299
1,70(59)xxx,xxCichlidaeCichlids nei101,805
1,70(59)052,xxSarotherodon, Tilapia sppTilapias nei278,837
1,74(03)001,01Thyrsites (= Leionura)atunsnoek57,669
1,74(04)xxx,xxTrichiuridaeHairtails, cutlassfishes119,796
1,74(04)003,02Trichiurus lepturusLargehead hairtail710,280
1,75(01)002,01Scomber japonicuschub mackerel1,739,889
1,75(01)002,05Scomber scombrusAtlantic mackerel600,556
1,75(01)014,xxRastrelliger sppIndian mackerels nei342,237
1,75(01)014,03Rastrelliger kanagurtaIndian mackerel176,824
1,75(01)015,xxScomberomorus sppSeerfishes nei68,551
1,75(01)015,03Scomberomorus commersonNarro-barred Spanish (king) mackerel114,316
1,75(01)015,12Scomberomorus niphoniusJapanes Spanish mackerel121,135
1,75(01)023,xxAuxis thazard, rocheiFrigate and bullet tunas146,794
1,75(01)024,06Euthnnus affinisKawakawa92,597
1,75(01)025,01Katsuwonus pelamisSkipjack tuna884,235
1,75(01)026,01Thunnus thynnusNorthern bluefin tuna36,993
1,75(01)026,05Thunnus alalungaAlbacore176,922
1,75(01)026,08Thunnus maccoyilSouthern bluefin tuna35,212
1,75(01)026,10Thunnus albacaresYellowfin tuna713,293
1,75(01)026,12Thunnus obesusBigeye tuna243,653
1,78(01)xxx,xxScorpaenidaeScorpionfishes, etc nei61,042
1,78(01)001,xxSebastes sppAtlantic redfishes400,680
1,78(07)014,02Pleurogrammus azonusAtka mackerel139,155
1,83(02)004,05Pleuronectes platessaEuropean plaice203,308
1,83(02)005,01Reinhardtius hippoglossoidesGreenland halibut80,236
1,83(02)014,01Hippoglossoides platessoidesAmerican plaice (= Long rough dab)80,911
1,83(02)024,02Limanda asperaYellowfin sole203,149
1,89(03)xxx,xxBalistidaeTriggerfishes, durgons39,746
1,89(03)004,xxCantherines (= Navodon) sppFilefishes272,979
1,89(03)010,01Stephanolepis cirrhiferThreadsail filefish256,529
2,28(01)001,xxPenaeus sppPenaeus shrimps nei412,879
2,28(01)001,01Penaeus aztecusNorthern brown shrimp70,858
2,28(01)001,03Penaeus merguiensisBanana prawn61,421
2,28(01)001,22Penaeus setiferusNorthern white shrimp44,596
2,28(04)002,03Pandalus borealisNorthern prawn193,191
2,28(07)009,03Acetes japonicusAkiami paste shrimp222,608
3,16(07)008,xxCrassostrea sppCupped oysters nei168,024
3,16(07)008,01Crassostrea gigasPacific cupped oyster562,866
3,16(07)008,03Crassostrea virginicaAmerican cupped oyster271,925
3,16(08)xxx,xxPectinidaeScallops nei75,191
3,16(08)003,07Pecten yessoensisJapanese scallop229,924
3,16(08)003,09Pecten maximusCommon scallop20,224
3,16(08)014,04Placopecten magellanicusSea scallop104,929
3,16(08)030,01Argopecten gibbusCalico scallop125,609
3,16(10)xxx,xxMytilidaeSea mussels nei135,454
3,16(10)001,05Mytilus edulisBlue mussel554,756
3,16(10)001,10Mytilus smaragdinusGreen mussel61,623
3,17(04)045,01Arctica islandicaOcean guahog195,173
3,17(10)020,01Spisula solidissimaSurf clam182,834
3,17(11)004,05Venus gallinaStriped venus27,348
3,17(11)018,01Venerupis japonicaJapanese(= Manila) clam166,134
3,17(11)041,xxPaphia sppShort neck clams86,060
3,17(11)075,01Mercenaria mercenariaHard clam32,460
3,17(21)025,02Corbicula japonicaJapanese clam53,873
3,21(02)xxx,xxSepia spp, Sepiola sppCuttlefishes, bobtail squids214,642
3,21(04)001,xxLoligo sppCommon squide119,570
3,21(05)010,03Illex argentinusArgentine shortfin squid43,706
3,21(05)058,03Todarodes pacificusJapanese flying squid192,466
3,21(09)xxx,xxOctopodiadeOctopuses210,102
3,21Loliginidae, OmmastrephidaeSquids nei852,553
3,99BivalviaClams nei648,774
6,17(41)007,xxRhopilema sppJellyfishes63,700

Source: FAO Yearbook of Fishery Statistics, Catches and Landings, Volume 64


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