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THE IMPORTANCE OF GENOTYPE, SIZE ON STOCKING AND STOCKING DATE TO THE SURVIVAL OF BROWN TROUT (Salmo trutta L.) RELEASED IN IRISH LAKES

M.F. O'Grady
Central Fisheries Board
Glasnevin, Dublin, Ireland

ABSTRACT

The relative success of a series of brown trout (Salmo trutta L.) stocking experiments in a range of Irish lakes are outlined. Data indicate that in the case of yearling plantings, the direct offspring of wild trout survived in greater numbers than an inbred strain of trout. Size on stocking can be more important than genotype in relation to plantings of two-year old fish. The larger individuals tend to survive in greater numbers and contribute more to angling catches. Repeated stockings of trout in spring and autumn indicate that the former are more successful both in terms of survival and subsequent angling catches.

RESUME

En Irlande, on a procédé, avec des résultats variables, à toute une série d'expériences de repeuplement des lacs en truites (Salmo trutta L.). On a constaté, en cas de repeuplement avec des sujets d'un an, que les descendants directs de truites sauvages avaient un taux de survie supérieure à celui d'une race consanguine de truites. Si l'on utilise des poissons de deux ans, leur taille au moment du repeuplement peut jouer un rôle plus important que le génotype. Les truites les plus grosses ont tendance à avoir un taux de survie supérieur et représentant une part plus importante des captures. On obtient de meilleurs résultats (taux de survie, captures) en repeuplant les eaux au printemps plutôt qu'en automne et donne des exemples de leurs effets sur les stocks de truites. Il passe en revue les problèmes qui subsistent et indique brièvement le travail qui reste à accomplir.

1. INTRODUCTION

The Central Fisheries Board (formerly the Inland Fisheries Trust) was founded in 1951 to develop and promote brown trout (Salmo trutta L.) and coarse fish (principally pike (Esox lucius L.), bream (Abramis brama L.), rudd (Scardinius erythrophthalmus L.) and tench (Tinca tinca L.)) sports fishing in the Republic of Ireland.

There are many lake brown trout fisheries in Ireland. The Central Fisheries Board developed these lakes by improving the trout spawning and nursery areas in inflowing streams and by controlling the trout's predators (pike) and competitors (perch, Perca fluviatilis (L.)) in the lakes.

In the late sixties a combination of arterial drainage schemes and increased instances of organic pollution in streams resulted in serious reductions in wild trout recruitment rates to lake fisheries. A mjaor stocking programme was undertaken at that time in many lakes to maintain optimum trout stocks. In 1975 a long-term research programme commenced to evaluate the overall development programme on trout lakes (O'Grady, 1981). The current manuscript outlines some of the results of this study - the relative importance of genotype, size on stocking and the actual stocking date to the survival of planted brown trout in a range of Irish lakes.

2. MATERIALS AND METHODS

2.1 The study areas

Experimental stockings were carried out in nine lakes. They ranged in size from 1 to 1 883 ha. Details of the size, depths, water chemistry, geology and resident fish stocks in these waters are presented (Table 1).

2.2 Trout genotypes

Five discrete brown trout strains were used in these experiments. The Central Fisheries Board have developed a fast-growing variety (Roscrea trout) over the past 25 years. A brood pool of 1 000 breeding pairs of Roscrea trout are retained annually. Seven different strains of wild trout have been crossbred with the brood pool at intervals to prevent excessive inbreeding. Details of the evolution of this stock are presented in O'Grady (1981). The four additional strains of trout used in these experiments were F' wild trout - the direct offspring of wild Irish lake trout which were reared in fish-farm conditions from the egg stage.

2.3 The collection of quantitative data on trout stocks

In four of the lakes, trout population estimates were carried out at intervals to assess the survival of fish stocked experimentally. A Petersen tag recapture type exercise was used. Tagged trout were stocked and the water was subsequently gillnetted for a three to five day period to obtain a sample of both the introduced tagged fish and the resident stock for estimate purposes. Data indicate that the standing crop estimates obtained in this way were accurate (O'Grady, 1981a, in press).

In the remaining waters, samples of trout from experimental stockings were recaptured in gillnet gangs which were capable of catching a random cross section of the population present (O'Grady, 1981a).

2.4 Recognition of specific trout stocks

All of the trout stocked during the course of these experiments were either fin-clipped or tagged for subsequent identification purposes. The tags utilized were Dennison floy anchor tags. Fin-clipped and tagged trout were never used in the same experiment in order to avoid the problem of possible differential mortalities from each marking technique. In experiments where fin clips were used for identification purposes and two genotypes were stocked, the same fin type was clipped in each group (i.e., a left and right pelvic or pectoral fin). Data indicate no differential mortality (P>0.05) in trout of identical genotype which has one or other pectoral or pelvic fin clipped (O'Grady, 1981).

3. RESULTS

3.1 The influence of genotype

In 1977 and 1978, seven relatively small lakes (0.4–130 ha) were stocked with autumn fingerlings or spring yearling trout. Each water was planted with fish of two strains, Roscrea trout and one F' wild group. Each group were marked by clipping a single fin. In October 1979, these waters were test-netted to establish the relative survival of the experimental stockings. Reasonable numbers of these fish were captured in four of the seven lakes. In all four waters there was a significantly higher (P < 0.05 or P < 0.001) survival of the F' wild strain (Table 2). In the remaining three lakes poor survival of both strains was probably caused by a combination of factors - predation by the resident pike stocks and both inter-and intra-species competition with resident trout and perch populations.

The Roscrea and F' wild trout used in these experiments were not homogeneous in terms of mean length values on stocking. The former group were larger in each case and maintained this differential (P<0.001) over the experimental period in at least three of the experiments (Table 3).

When the samples of fish recaptured in each lake were being removed from the gillnets, it was noted that some of the immature fish were soft-fleshed. Subsequently, when individual fish were being weighed and measured, the flesh texture of each immature trout was recorded. A distinct pattern emerged. All of the immature Roscrea trout were soft-fleshed. Their F' wild counterparts had much firmer musculature. Trout which would have matured later in the year were not utilized in this comparative test because maturing females tend to become soft-fleshed and ripening males usually develop a toughened skin.

3.2 The importance of size on stocking

The relevance of size on stocking to the subsequent survival of planted trout was illustrated in a number of experimental programmes. In each case, the larger group of fish survived and/or were cropped by anglers in greater numbers. In each experiment, the larger group of fish were significantly (P<0.001) bigger than their counterparts.

A. Roscrea two-year old and yearling trout

Both Roscrea two-year old and yearling fish were planted in Lough O'Flyn in April 1976. A large percentage (56.4 percent) of the former group were cropped by anglers in 1976 (April to September) and a significant number of these fish (33.1 percent) were still present in the fishery in October 1976. An additional 12 and 6 percent of this two-year old group were caught by anglers during the 1977 and 1978 angling seasons respectively. In contrast, none of the yearling Roscrea fish were recorded in angling catches in 1977 or in subsequent years. Five trout population estimates carried out in Lough O'Flyn at intervals from October 1976 to October 1979 indicate that few of these fish has survived (<3.0 percent) 17 months after planting.

A similar stocking programme in Lough Inchiquin yielded identical results with a greater number of the two-year old Roscrea fish surviving for a longer period (Tables 4 and 5).

B. Roscrea and F'Owel two-year old trout

Roscrea and F'Owel two-year old trout were stocked concurrently on three occasions. In one instance (O'Flyn, 1977) F'Owel trout were larger than their Roscrea counterparts. In the other two experiments (O'Flyn, 1976 and Inchiquin, 1976) the Roscrea trout were the larger fish. In each of the three experiments the larger batch of trout, irrespective of genotype, survived and/or were caught in greater numbers by anglers (Tables 4 and 5).

3.3 The influence of stocking date

Repeated planting of autumn yearling (19 months old) and two-year old (22–24 months old) Roscrea trout in Loughs O'Flyn and Sheelin illustrate the importance of the stocking date to the subsequent survival of planted trout. In each case, the two-year old fish survived in greater numbers for a longer period and thus more of these fish were cropped by anglers (Tables 6 and 7).

The numbers of autumn yearling trout which survived this first overwintering period, post-stocking, were checked in relation to four specific plantings (Sheelin, October, 1977 and 1978; O'Flyn, October 1976 and 1977). In both the Lough Sheelin stockings and in the O'Flyn October 1976 planting, only a small (8.9–25.6 percent) proportion of each stocked group survived overwinter - a 5–6 month period.

The 95 percent confidence limits obtained for the estimated number of autumn yearlings (October 1977) still present in Lough O'Flyn in February 1978 were too broad to indicate the precise overwintering mortality for this group. However, very few of these trout were caught by anglers in 1978 and 1979 (9.9 and 10.3 percent) and none of these fish were captured two years after planting during a population estimate (Tables 6 and 7). These data indicate a poor survival rate for this stocking.

4. DISCUSSION

The results of these experiments indicate that genotype can influence the survival of stocked trout. Significantly greater numbers of F' yearling trout survived than their Roscrea counterparts in four small lakes. A similar pattern was noted for domesticated and F' wild rainbow trout (Borgeson, 1966; Webster and Flick, 1974 and Reisenbichler and McIntyre, 1977) and brook trout (Flick and Webster, 1962 and Mason et al., 1967). The same trend has also been observed between different domesticated trout strains. (Cordone and Nicola, 1970: Brown, 1970; Rawstron, 1973 and Ayles, 1974) and also between transplanted wild and domesticated brown trout (Miller, 1953 and Strange and Kennedy, 1979).

The higher survival of F' genotypes in this study might be related to the fact that they were introduced into waters, similar ecologically to those occupied by their parental stocks. Ayles (1974) and Webster and Flick (1974) have stressed the importance of matching planted strains to specific habitats. The F' genotypes may also have higher stamina than their Roscrea counterparts. Green (1964) noted that F' brook trout have higher stamina than domesticated varieties. Fishfarm experiments indicated that Roscrea fish were tamer than the F' strains (O'Grady, 1981), a factor which might render the former group more prone to predation. Greene (1951), Vincent (1960) and Butler (1974) quoting McClaren also record that domesticated trout display a lack of wariness even after release from fish farms. Fishfarm experiments also indicated that F' genotypes will adopt a territorial stance given the opportunity (O'Grady, 1981).

Roscrea fish have difficulty in recognizing invertebrate food items for at least ten days after release and do not feed at the same rate as resident fish during this period (O'Grady, 1981). F' wild trout may not experience this difficulty. A combination, of some, or all, of these factors probably resulted in the higher survival of the F' genotypes. The unusually soft musculature of the Roscrea fish which did survive may also reflect the social stress which these fish were subjected to in an alien and competitive environment.

Size on stocking is of critical importance to survival and also influences the number of fish caught by anglers. The importance of this factor in relation to yearling and 2+ fish has been recognized previously for both brown trout (Brown, 1970; Cooper, 1974 and Fleming-Jones, 1978) and rainbow trout (Borgeson, 1966; Brown, 1970; Cooper, 1974 and Fleming-Jones, 1978). In Loughs O'Flyn and Inchiquin size on release was more important than genotype in determining subsequent angling returns. The cropping of larger trout in two-year old plantings has been noted previously (Shetter, 1944; Needham, 1947; Larson and Ward, 1955; Mullan, 1956; Hallock et al., 1961; Wagner and Wallace, 1963; Hansen and Stauffer, 1971; Templeton, 1971 and Cooper, 1974). The higher returns to anglers of larger trout might be related to an increased ability of these individuals to survive in competitive situations.

The date of planting can, of itself, seriously influence survival. Repeated autumn yearling plantings in Loughs O'Flyn and Sheelin failed to overwinter successfully. Comparable two-year old spring stockings survived in much greater numbers. Many authors report a similar differential survival pattern for autumn and spring stockings of non-migratory salmonids (Shetter and Hazzard, 1940; Needham and Slater, 1944; Shetter, 1944; Reimers, 1957; Buettner, 1961; Anderson, 1962; Brown, 1970; Templeton, 1971 and Cooper, 1974). A paucity of food in the autumn has been suggested as the reason for the poor survival (Brown, 1970). This does not hold in Irish lakes because autumnal standing crops for many invertebrate species are relatively high (Whelan and O'Connor, pers.comm.). Data indicate that, initially in Irish waters stocked fish tend to be surface feeders, ignoring the benthic invertebrates (O'Grady, 1981). Thus, an absence of surface food items in the autumn may result in the poor survival of fish stocked at this time of year.

The results of these experiments illustrate that several factors, genotype, size on planting and the stocking date, can seriously influence survival and therefore affect angling catches. These data will allow a rationalization of stocking procedure in Irish lake brown trout fisheries and contribute to a more efficient fisheries management programme. There are many small (1–30 ha) productive lakes in Ireland which are not fished intensively. Annual spring plantings of yearling F' genotype fish in these waters will ensure the maintenance of high stocks in circumstances where natural production is limited or absent. In larger waters, annual spring plantings of large two-year old trout will successfully supplement natural stocks. A 50 percent reduction of stock densities in Central Fisheries Board fish farms since 1978 has resulted in an increased growth rate of stocks (O'Grady, 1981). Two-year old Roscrea trout now (1981) average 30 cm in length. The Roscrea strain fish are the most useful for stocking purposes where two-year old trout are required because they grow more quickly in fishfarm conditions than F' strain fish (O'Grady, 1981). The reduction in fishfarm production because of the decrease in pond stock densities has been offset by the adoption of a cage-culture system in lakes.

5. ACKNOWLEDGEMENTS

I am very grateful to the Central Board for providing me with the facilities to carry out this research. Special thanks are due to T. Sullivan and P. Green for technical assistance. I wish to gratefully acknowledge all assistance received from members of both the development and fishfarm staffs of the Central Fisheries Board. Sincere thanks are due to Drs. K. Whelan and J. O'Connor for information on invertebrate standing crops in Irish lakes.

6. REFERENCES

Anderson, R.B., 1962 A comparison of returns from fall- and spring-stocked hatchery-reared lake trout in Maine. Trans.Am.Fish.Soc., 91:425–7

Ayles, G.B., 1975 Influence of the genotype and the environment on growth and survival of rainbow trout (Salmo gairdneri) in central Canadian aquaculture lakes. Aquaculture, 6:181–8

Borgeson, D.P., 1966 Trout lake management. In Inland fisheries management, edited by A. Cahoun. Department of Fish and Game, Sacramento, California. pp. 168–78

Brown, K.S., 1970 Reservoir trout fishing. Salm.Trout.Mag., 188:33–5

Buettner, H.J., 1961 Recoveries of tagged, hatchery-reared lake trout from Lake Superior. Trans.Am.Fish.Soc., 90:404–12

Butler, R.L., 1974 Some thoughts on the effects of stocking hatchery trout on wild trout populations. In Proceedings of Wild Trout Management Symposium at Yellowstone National Park. pp. 40–7

Cooper, E.L., 1974 Trout in streams. Trout, 15(1):4–11

Cordone, A.J. and S.J. Nicola, 1970 Harvest of four strains of rainbow trout Salmo giardneri, from Beardsley Reservoir, California. Calif.Fish Game, 56(4):271–87

Fleming-Jones, D., 1978 Recreational freshwater fisheries, their conservation, management and development (1977). Oxford, Keble College, pp. 36–48

Flick, W.A. and D.A. Webster, 1962 Problems in sampling wild and domestic stocks of brook trout (Salvelinus fontinalis). Trans.Am.Fish.Soc., 91:140–4

Greene, C.W., 1951 Results from stocking brown trout of wild and hatchery strains at Stillwater Pond. Trans.Am.Fish.Soc., 81:43–52

Greene, D.M., Jr., 1964 A comparison of stamina of brook trout from wild and domesticated parents. Trans.Am.Fish.Soc., 93:96–100

Hallock, R.J., W.F. Van Woert and L. Shapovalov, 1964 An evaluation of stocking hatchery-reared steelhead rainbow trout (Salmo gairdneri) in the Sacramento River system. Fish.Bull.Calif.Dep.Fish Game, (114):74 p.

Hansen, M.J. and T.M. Stauffer, 1971 Comparative recovery to the creel, movement and growth of rainbow trout stocked in the Great Lakes. Trans.Am.Fish.Soc., 100(2):336–49

Larson, R.W. and J.M. Ward, 1955 Management of steelhead trout in the State of Washington. Trans.Am.Fish.Soc., 84:261–74

Mason, J.W., O.M. Brynildon and P.E. Degurse, 1967 Comparative survival of wild and domestic strains of brook trout in streams. Trans.Am.Fish.Soc., 96:313–9

Miller, R.B., 1953 Comparative survival of wild and hatchery-reared cutthroat trout in a stream. Trans.Am.Fish.Soc., 83:120–30

Mullan, J.W., 1956 The comparative returns of various sizes of trout stocked in Massachusetts streams. Prog.Fish-Cult., 18(1):35 p.

Needham, P.R., 1947 Survival of trout in streams. Trans.Am.Fish.Soc., 77:26–31

Needham, P.R., and D.W. Slater, 1944 Survival of hatchery-reared brown and rainbow trout as affected by wild trout populations. J.Wildl.Manage., 8:22–36

O'Grady, M.F., 1981 A study of brown trout (Salmo trutta L.) populations in selected Irish lakes. Thesis submitted in candidature for the Ph.D. degree to the National University of Ireland.

O'Grady, M.F., 1981a Some direct gillnet selectivity tests for brown trout populations. Ir.Fish.Invest.Ser.A., (22):9 p.

O'Grady, M.F., The development of a lake trout (Salmo trutta L.) population estimate technique suitable for Irish waters. Ir.Fish.Invest.Ser.A., (in press)

Rawstron, R.R., 1973 Comparisons of disk dangler, trailer and internal anchor tags on three species of salmonids. Calif.Fish.Game, 59(4):266–80

Reimers, N., 1957 Some aspects of the relation between stream foods and trout survival. Calif.Fish.Game, 34:43–69

Reisenbichler, R.R. and J.D. McIntyre, 1977 Genetic differences in growth and survival of juvenile hatchery and wild steelhead trout Salmo gairdneri. J.Fish.Res.Board Can., 34(1):123–8

Shetter, D.S., 1944 Further results from spring and fall plantings of legal-sized hatchery-reared trout in streams and lakes of Michigan. Trans.Am.Fish.Soc., 74:35–57

Shetter, D.S. and A.S. Hazzard, 1940 Results from plantings of marked trout of legal size in streams and lakes of Michigan. Trans.Am.Fish.Soc., 70:446–68

Strange, C.D. and C.J.A. Kennedy, 1979 Yield to anglers of spring and autumn stocked hatchery reared and wild brown trout (Salmo trutta L.). J.Inst.Fish.Manage., 10(2):45–51

Templeton, R.G., 1971 An investigation of the advantages of autumn and spring stocking with brown trout in a Yorkshire reservoir. J.Fish Biol., 3:303–24

Vincent, R.E., 1960 Some influences of domestication upon three stocks of brook trout. Trans.Am.Fish.Soc., 89:35–52

Wagner, H.H. and R.L. Wallace, 1963 The seaward migration and return of hatchery-reared steelhead trout Salmo gairdneri Richardson, in the Alser River, Oregon. Trans.Am.Fish.Soc., 92(3):202–10

Webster, D.A. and W.A. Flick, 1974 Species management. In Proceedings of the Wild Trout Management Symposium at Yellowstone National Park, pp. 40–7

Table 1 Details of the study areas

LakeArea
(ha)
Max.depth
(m)
Mean depth 
(m)
AlkalinityConductivity Solid geologyFish species
Sheelin1 8831452.70287Carboniferous limestoneBt, Pe, Pi, Ro, Ru, Br, T, St(3),St(9)
Mount Dalton8435.00544Carboniferous limestoneBt
Doolin Pond2.22.71.3--Carboniferous limestoneBt, St(3), C
O'Flyn130194.52.80282Carboniferous limestoneBt, Pe, Pi, Ru, Br, St(3), St(9)
Inchiquin1132763.70400Carboniferous limestone sandstone, millstone grit, flagstone, shalesBt, Pe, Pi, Ru, T, St(3), St(9)
Muckanagh3002034.00405Carboniferous limestoneBt, Pe, Pi, Ru, Br
Kilkee Reservoir2.43.61.80.85300Millstone grit, flagstoneBt
Gortglass311530.,50104Millstone grit, flagstoneBt. Ch
Gloster Pond1----Carboniferous limestoneBt, Pe, Pi, Ru

Bt = brown trout;
Pe = perch;
Pi = pike;
Ru = rudd;
Ro = roach;
Br = bream;
T = tench;
St(3) = three-spined stickleback;
St(9) = nine-spined stickleback;
C = carp;
Ch = char

Eels present in all lakes

Alkalinity as milli equivalents per litre

Conductivity as reciprocal megohms

Table 2 The comparative survival of Roscrea and F' wild trout strains, stocked in a number of small lakes, as indicated by x2 tests

LakeStrainNumber stocked
(1977/78)
Number recaptured
(1979)
X2
Significance
GortglassRoscrea8294  
Mask F'8293927.8
P<0.001***Mask F'>Roscrea
Mount DaltonRoscrea1 00023  
Mask F'1 000383.6
P<0.05* Mask F'>Roscrea
Doolin PondRoscrea60014  
Sheelin F'60010462.8
P<0.001*** Sheelin F'>Roscrea
Kilkee ReservoirRoscrea50023  
Sheelin F'500383.5
P<0.05 * Sheelin F'>Roscrea
O'FlynRoscrea4 0083  
O'Flyn F'4 0085-
Insufficient data
MuckanaghRoscrea3 1750  
Inchiquin F'3 1750--
Gloster PondRoscrea5000  
Sheelin F'5000--

* = significant difference;

*** = very highly significant difference

Table 3 A statistical comparison of the mean (x) length values of Roscrea and F' wild trout on stocking and subsequently on recapture in the Mount Dalton Lake, Kilkee Reservoir and Doolin Pond experiments

  On stocking (November 1977) On recapture (October 1979)
LakeTrout strainMean(x) length
cm)
SEt valueSig.SampleMean(x) length
(cm)
SEt valueSig.Sample
Mount
Dalton
Roscrea13.90.316.74
P<0.001 ***
50 fish36.10.416.20
P<0.001 ***
23 fish
Mask F'11.50.2950 fish33.50.3738 fish
Kilkee ReservoirRoscrea13.60.286.10
P<0.001 ***
50 fish41.50.404.26
P<0.001 ***
23 fish
Sheelin F'11.20.2450 fish38.90.3838 fish
Doolin
Pond
Roscrea12.50.273.20
P<0.001 ***
50 fish30.10.364.03
P<0.001 ***
14 fish
Sheelin F'11.60.3050 fish28.40.3150 fish

*** = Very highly significant difference

SE = Standard error of the mean

Table 4 Data illustrating the importance of size to the survival of stocked trout

LakeStrainAge
(months)
X Length
(S.E.)
No. Fish StockedDateStanding Crop Estimates ()
       Oct '76
Feb '77Oct '77Feb '78Oct '79
O'FlynRoscrea2428.3(0.13)3,323April '76(1,262-694)
1,100
(1,395-586)
960
(209-10)
201
(332-20)
211
0
 Roscrea1314.7(0.13)8,735May '76(280-47)
235
(265-1)
221
(254-1)
206
(247-1)
204
0
       Aug. '76Aug '77Sept '78  
InchiquinRoscrea2419.8(0.27)2,806April '76(1,000-233)
617
00  
 Roscrea12 14.5(0.21)6,313April '76000  
       Oct '76Feb '77Oct '77Feb '78Oct '79
O'FlynRoscrea2221.4(0.13)850Feb '77--(354-32)
196
(396-87)
203
 0
 F'Owel2223.4(0.16)1,175Feb '77--(566-116)
364
(883-141)
568
0
O'FlynRoscrea2228.2(0.13)3,323April '76(1,262-694)
1,100
(1,395-586)
960
(209-1)
201
(332-20)
211
0
 F'Owel2219.8(0.13)4,612April '76(892-580)
784
(1,247-612)
1,199
(640-280)
530
(690-261)
533
(70-1)
50
       Aug. '76Aug. '77Sept '78  
InchiquinRoscrea2419.8(0.27)2,806April '76(1,000-233)
617
00  
 F'Owel2417.3(0.32)3,300April '76(666-93)
380
(386-1)
197
0  

(S.E.) - Standard error of the mean length on stocking;
(-) - 95% Confidence limits.

Table 5 Data illustrating the importance of size in relation to angling catches of stocked trout

LakeStrainAge
(months)
X Length(S.E.)No. Fish StockedDateAngling Catch Estimates ()
       19761977197819791980
O'FlynRoscrea2428.3(0.13)3,323April '76(2,137-1,740)
1,876
(419-379)
399
(209-87)
199
00
Roscrea1314.7(0.13)8,735May '7600000
InchiquinRoscrea2419.8(0.27)2,806April '76(1,629-1,132)
1,352
(322-1) 159000
Roscrea1214.5(0.21)6,313April '7600000
O'FlynRoscrea2221.4(0.13)850Feb '77-(86-3)
480
000
F'Owel2223.4(0.16)1,175Feb '77-(437-128) 294(440-101)
320
00
O'FlynRoscrea2228.2(0.13)3,323April '76(2,137-1,740)
1,876
(419-379) 399(209-87)
199
00
F'Owel2219.8(0.13)4,612April '76(1,422-1,325)
1,384
(401-278) 387(331-74)
270
00
InchiquinRoscrea2419.8(0.27)2,806April '76(1,629-1,132)
1,352
(322-1)
159
000
F'Owel2417.3(0.32)3,300April '76(617-273)
462
(321-0)
155
000

(S.E.) - Standard error of the mean length on stocking;
(-) - 95% confidence limits.

Table 6 Data illustrating the importance of the stocking date to the survival of planted brown trout

LakeStrainAge
(months)
X Length (S.E.)No.Fish StockedDateStanding Crop Estimates ()
       March'78March'79   
SheelinRoscrea2322.4(0.22)33,296March'77(21,723-10,699)
16,211
(13,937-6,163)
10,281
   
Roscrea1921.3(0.26)20,827Oct '77(4,897-1,471)
3,184
(5,956-3,971)
4,972
   
SheelinRoscrea2326.3(0.18)34,133March'78-(8,180-5,454)
6,820
   
Roscrea1925.7(0.2)15,200Oct '78-(4,685-3,123)
3,900
   
       Oct'76Feb '77Oct '77Feb '78Oct '79
O'FlynRoscrea1924.3(0.14)3,338Oct '75(331-21)
163
(460-130)
280   
(290-1)
131
(285-1)
127
 0
Roscrea2428.2(0.14)3,323April '76(1,262-694)
1,100
(1,395-586)
960
(209-1)
201
(332-20)
211
 0
O'FlynRoscrea1921.5(0.15)3,978Oct '76-(490-260)
354
(560-73)
303
(530-71)
301
(47-1)
20
Roscrea2221.4(0.13)850Feb '77--(354-32)
196
(396-87)
203
0
O'FlynRoscrea1921.5(0.15)1,265Oct '77---(1,202-291)
759
 0
Roscrea2222.3(0.17)1,081Feb '78----(510-112)
216

(S.E.)- Standard error of the mean length on stocking;
(-) - 95% confidence limits.

Table 7 Data illustrating the importance of the stocking date in relation to angling catches of stocked trout

LakeStrainAge
(months)
X Length(S.E.)No. Fish
Stocked
DateAngling Catch Estimate ()
       197719781979  
SheelinRoscrea2322.4(0.22)33,296March '77(*)(1,841-1,287) 1,584(1,423-364) 814  
Roscrea1921.3(0.26)20,827Oct'77(*)(*)(190-0) 28  
SheelinRoscrea2326.3(0.18)34,133March '78-(*)(1,315-211) 701  
Roscrea1925.7(0.2)15,200Oct '78--0  
       19761977197819791980
O'FlynRoscrea1924.3(0.14)3,338Oct'75(390-110)
294
(108-30)
68
000
Roscrea2428.2(0.14)3,323April'76(2,137-1,740) 1,876(419-379)
399
(209-87) 19900
O'FlynRoscrea1921.5(0.15)3,978Oct'76-(251-90)
163
(230-43) 141(135-22) 930
Roscrea2221.4(0.13)850Feb'77-(86-3)
48
000
O'FlynRoscrea1921.5(0.15)1,265Oct'77--(241-21) 125(165-95) 1300
Roscrea2222.3(0.17)1,081Feb'78--(490-141) 224(460-100) 227(170-30) 113

(*) - Trout which were below the angling size limit and not cropped;
(S.E.) - Standard error of the mean length on stocking;
(-) - 95% confidence limits.


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