Animal health

Emerging zoonotic coronaviruses in animals situation update

25 September 2025, 17:00 hours; Rome

Overview

Situation: Emerging zoonotic coronaviruses pose a persistent threat to global health owing to their ability to cross species barriers and cause unpredictable outbreaks presenting an ongoing pandemic risk. This FAO situation update focusses on: (1) Severe Acute Respiratory Syndrome Coronavirus-2 (SARS-CoV-2): a virus that is spreading globally through human-to-human transmission but has also demonstrated ability to infect multiple animal species, and (2) Middle East Respiratory Syndrome Coronavirus (MERS-CoV): zoonotic virus with pandemic potential. The list of animal families in which at least one species tested positive for the virus RNA are presented in Table 1.

SARS-CoV-2MERS-CoV

Bovidae, Canidae, Cebidae, Cercopithecidae, Cervidae, Cricetidae,* Felidae, Hominidae, Hyaenidae, Mustelidae,* Procionidae, Viverridae, Hippopotamidae, Myrmecophagidae, Atelidae, Rhinocerotidae, Suidae, Agamidae, Phasianidae, Anatidae, Castoridae, Muridae, Chlamyphoridae, Leporidae, Vespertilionidae, Sciuridae, Didelphidae, Procyonidae and Fringillidae

Camelidae,** Bovidae, Equidae, Emballonuridae, Ixodidae

* In rare occasions, spill-back of SARS-CoV-2 from animals to humans has been evidenced in family Mustelidae (mink-to-human in the Netherlands [reference] and in Denmark [reference]; and family Cricetidae (hamster-to-human in Hong Kong Special Administrative Region (China) [reference]).
** MERS-CoV spillovers originating from species of the Camelidae family (Dromedary camel-to-human) have been observed in many zoonotic events in the Arabian Peninsula.

 

Reported human cases: The new and cumulative number of human cases as of 31 August 2025 is presented in Table 2.For detailed information on MERS-CoV human cases, please refer to WHO MERS-CoV information page, and for COVID-19 human cases, please refer to WHO COVID-19 Dashboard and WHO COVID-19 Weekly Updates.

 SARS--2MERS-CoV

Total confirmed human cases

778 572 430

2 628

Total number of fatal cases

7 101 070

948

New confirmed cases since 1 June 2025

522 255

2*

New deaths since1 June 2025

4 135

1*

* The new MERS-CoV human cases were reported in the Kingdom of Saudi Arabia.

 

Situation in animals

As of 25 September 2025, SARS-CoV-2 has been detected in 68 animal species across 49 countries and territories, while MERS-CoV has been identified in 7 animal species across 16 countries.

Countries and territories that reported any virological findings in animals are outlined in Table 3 and Map 1, while Table 4 lists the animal species known to have been naturally infected by SARS-CoV-2 or MERS-CoV. The susceptibility of animal species for SARS-CoV-2 or MERS-CoV infection in experimental settings is detailed in Table 5.

RegionSARS-CoV-2MERS-CoV
Americas Argentina, Brazil, Chile, Ecuador, Mexico, Peru, Puerto Rico, Uruguay, Canada, Colombia, United States of America-

Asia and Pacific

China (Hong Kong SAR), India, Indonesia, Japan, Mongolia, Myanmar, Republic of Korea, Singapore, Thailand, Viet Nam

-

North Africa and Near East

Egypt

Egypt, United Arab Emirates, Iran, Iraq, Jordan, Kuwait, Morocco, Oman, Qatar, Saudi Arabia, Tunisia

Sub-Saharan Africa

Nigeria, Senegal, South Africa

Burkina Faso, Ethiopia, Kenya, Nigeria, Senegal

Europe

Belgium, Bosnia and Herzegovina, Bulgaria, Croatia, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Netherlands (Kingdom of the), Poland, Portugal, Slovenia, Spain, Sweden, Switzerland, United Kingdom of Great Britain and Northern Ireland, Russian Federation

-

* in alphabetical order. Countries and territories that published virological findings in animals since the last update are highlighted orange. Include animal cases officially reported by national authorities and the WOAH , or positive virological findings referred to in scientific publications . Please see the respective articles of the recent additions under section “recent publications”.

 

Map 1. SARS-CoV-2 and MERS-CoV events* in animals published or reported at national level, as of 25 September 2025

* Events include animal cases officially reported by national authorities and the WOAH, or positive virological findings referred to in scientific publications.

Notes: Refer to the disclaimer available on this webpage for the names and boundaries in this map. Final boundary between the Sudan and South Sudan has not yet been determined. Final status of the Abyei area is not yet determined. Dotted line represents approximately the Line of Control in Jammu and Kashmir agreed upon by India and Pakistan. The final status of Jammu and Kashmir has not yet been agreed upon by the parties.
Source: United Nations Geospatial. 2020. Map of the World. [Cited September 2025]. Modified with data from WOAH and scientific publications, 2025.

SARS-CoV-2MERS-CoV

Domestic cat

Felis catus

Dromedary camel

Camelus dromedarius

Domestic Dog

Canis lupus familiaris

Domestic Donkey

Equus africanus asinus

Domestic American Mink

Neovison vison

Domestic Cattle

Unspecified

Domestic Ferret

Mustela furo

Domestic Sheep

Ovis aries

Wild American Mink

Neovison vison

Domestic Goat

Unspecified

Western lowland Gorilla

Gorilla gorilla gorilla

Egyptian tomb bat

Taphozous perforatus

White-tailed deer

Odocoileus virginianus

Camel tick

Hyalomma dromedarii 

Binturong

Arctictis binturong

 

 

Coatimundi

Nasua nasua

 

 

Fishing cat

Prionailurus viverrinus

 

 

Tiger

Panthera tigris

 

 

Lion

Panthera leo

 

 

Puma

Puma concolor

 

 

Snow Leopard

Panthera uncia

 

 

Indian Leopard

Panthera pardus fusca

 

 

Canada Lynx

Lynx canadensis

 

 

Spotted hyenas

Crocuta crocuta

 

 

Asian small-clawed otters

Aonyx cinereus

 

 

Hamster

Unspecified

 

 

Wild Eurasian River Otter

Lutra lutra

 

 

Hippopotamus

Hippopotamus amphibius
Unspecified

 

 

Black-Tailed Marmoset

Mico melanurus

 

 

Mule deer

Odocoileus hemionus

 

 

Antillean manatees

Trichechus manatus manatus

 

 

Giant anteater

Myrmecophaga tridactyla

 

 

Mandrill

Mandrillus sphinx

 

 

Monkey Squirrel

Saimiri sciureus

 

 

Red fox

Vulpes vulpes

 

 

Cattle

Unspecified

 

 

Buffalo

Unspecified

 

 

Goat

Unspecified
Capra hircus coreanae

 

 

Black- and brown headed Spider Monkey

Ateles fusciceps

 

 

Common woolly monkey

Lagothrix lagothricha

 

 

White rhinoceros

Ceratotherium simum

 

 

Duck a

Unspecified

 

 

Chicken a

Unspecified

 

 

Turkey a

Unspecified

 

 

Sheep

Unspecified

 

 

Pig

Unspecified

 

 

Lizard

Agama agama

 

 

Eurasian beaver 

Castor fiber

 

 

White-fronted capuchin

Cebus unicolor

 

 

House mouse

Mus musculus

 

 

Brown rat

Rattus norvegicus

 

 

Big hairy armadillo

Chaetophractus villosus

 

 

Pantanal cat  

Leopardus braccatus

 

 

Gray brocket

Subulo gouazoubira

 

 

Red deer

Cervus elaphus

 

 

Manned wolf

Chrysocyon brachyurus

 

 

European fallow deer

Dama dama

 

 

Eastern deer mouse

Peromyscus maniculatus

 

 

Raccoon

Procyon lotor

 

 

Eastern cottontail 

Sylvilagus floridanus

 

 

Eastern red bat

Lasiurus borealis

 

 

Groundhog

Marmota monax

 

 

Virginia opossum

Didelphis virginiana

 

 

Manned wolf

Ateles fusciceps

 

 

European fallow deer

Lagothrix lagothricha

 

 

Eastern deer mouse

Ceratotherium simum

 

 

Raccoon

Unspecified

 

 

Eastern cottontail 

Unspecified

 

 

Eastern red bat

Unspecified

 

 

Groundhog

Unspecified

 

 

Virginia opossum

Unspecified

 

 

White-eared opossum

Didelphis albiventris

 

 

European polecat

Mustela putorius

  

Cactus mouse

Peromyscus eremicus

  

House finch

Haemorhous mexicanus

  

Source: WOAH WAHIS, country reports and peer-reviewed journals3. Please see the respective articles of the recent additions under section “recent publications”.
3 Information from preprints is not included in this table.

 

Animal speciesScientific name
(wild animals)
Susceptibility
SARS-CoV-2MERS-CoV

Raccoon dogs

Nyctereutes procyonoides

Yes (reference)

 

Red Fox

Vulpes vulpes

Yes (reference)

 

Coyotes

Canis latrans

No (reference)

 

Deer mice

Peromyscus maniculatus

Yes (reference)

 

Bank voles

Myodes glareolus

Yes (reference)

 

Bushy-tailed woodrats

Neotoma cinerea

Yes (reference)

 

Laboratory BALB/c mice

 

Yes (reference)

 

White-tailed deer

Odocoileus virginianus

Yes (reference)

 

Ferret

Mustela furo

Yes (reference)

No (reference)

Egyptian fruit bat

Rousettus aegyptiacus

Yes (reference)

 

Striped skunks

Mephitis mephitis

Yes (reference)

 

Zebra fish

Danio rerio

Yes (reference)

 

Zebra mussel

Dreissena polymorpha

Yes (reference1) (reference2)

 

Syrian hamsters

Mesocricetus auratus

Yes

No (reference)

Tree shrews

Tupaia belangeri chinensis

Yes (reference1) (reference2)

 

Rhesus macaques

Macaca mulatta

Yes (reference)

Yes (reference1) (reference2)

The crab-eating macaque

Macaca fascicularis

Yes (reference)

 

Baboons

Papio hamadryas

Yes (reference)

 

Common marmosets

Callithrix jacchus

Yes (reference)

Yes (reference1) (reference2)

Cynomolgus macaques

Macaca fascicularis

Yes (reference)

 

African green monkeys

Chlorocebus aethiops

Not susceptible (reference)

Yes (reference)

Mosquitoes

Aedes aegypti, Aedes. albopictus, Culex tarsalis and Culex quinquefasciatus

Not susceptible (reference1) (reference2)

 

Midge

Culicoides sonorensis

Not susceptible (reference)

 

Chicken – Duck – Geese – Turkey – Quail and Pigeon

Unspecified

Not susceptible (reference)

 

Pig

Unspecified

Yes, Low susceptibility (reference1) (reference2)
(reference3)

Yes, low susceptibility (reference)

Cattle

Unspecified

Yes, Low susceptibility  (reference1) (reference2) (reference3)

 

Horse

Unspecified

No (reference)

Yes, low susceptibility (reference)

Sheep

Unspecified

Yes, Low susceptibility (reference)

Yes, low susceptibility (reference)

Goat

Unspecified

Yes,Low susceptibility (reference1) (reference2)

Yes, low susceptibility (reference)

Alpaca

Unspecified

No (reference)

Yes (reference)

Rabbit

Unspecified

Yes (reference)

Yes (reference)

Cat

Unspecified

Yes (reference)

 

Dog

Unspecified

Yes, Low susceptibility (reference)

 

Sprague Dawley rats

Rattus norvegicus

Yes (reference)

 

Elk

Cervus canadensis

Yes (ancestral virus),  
Low susceptibility to Delta VOC,  (reference1) ( reference2)

 

Mule deer

Odocoileus hemionus

Yes (reference)

 

Mexican free-tailed bats

Tadarida brasiliensis

Yes (reference)

 

Little brown bats  

Myotis lucifugus

No (reference)

 

Llama

Llama glama

 

Yes (reference)

Bactrian camel

Camel bacterinus

 

Yes (reference)

Jamaican fruit bat 

Artibeus jamaicensis

 

Yes (reference)

Information in this table should be interpreted with caution. Susceptibility of a given species to SARS-CoV-2 and MERS-CoV can vary depending on experimental conditions, including the virus strain, viral dose, inoculation method, and other factors.

 

Recent publications

Barry, M. (2025). [reference]. Epidemiological Characteristics of MERS‑CoV Human Cases, 2012‑2025. Journal of Epidemiology and Global Health, 15, Article 103.

Chen, C., Nooruzzaman, M., Varga, C., Naru, A., Cohen, H. R., Varvil, M. S., Hernandez, N., Diel, D. G., Fang, Y., & others. (2025). [reference]. Evaluating the distribution and clustering of SARS-CoV-2 antibodies in dogs across the United States of America. Scientific Reports, 15, 21758.

Choi, A., Stout, A. E., Rollins, A., Wang, K., Guo, Q., Jaimes, J. A., Kennedy, M., Wagner, B., & Whittaker, G. R. (2025). [reference]. SARS-COV-2 serosurvey of healthy, privately owned cats presenting to a New York City animal hospital in the early phase of the COVID-19 pandemic (2020-2021). In press, IJID One Health.

Hassan, A. M., Yasir, M., Al-Subhi, T., Sohrab, S. S., Abbas, A. T., El-Kafrawy, S. A., Kumosani, T., & Azhar, E. I. (2025). [reference]. Analysis of the metagenomic assembled genome of MERS-CoV and Alpha-CoV from Camelus dromedarius in Saudi Arabia. Journal of infection and public health, 18(11), 102908. Advance online publication.

Karani, A., Ngere, I., Ombok, C., Singh, D., Jaoko, W., Njenga, M. K., & Palmer, G. H. (2025, June 17). [reference]. Low prevalence of Middle East Respiratory Syndrome Coronavirus infection in camel-exposed patients presenting with respiratory symptoms in Northern Kenya. The American Journal of Tropical Medicine and Hygiene. Advance online publication.

Kuhlmann, P. A., Di Nucci, D. L., Valdez, S. N., Marfia, J. I., Smith, I., Miranda, M. V., Fernandez, C. S., Grassi, E., Ruuth, E., & Salvatierra, K. A. (2025). [reference]. SARS-CoV-2 Antibodies in Wild Mammals from Atlantic Forest Region in Misiones province, Argentina. J Wildl Dis. Advance online publication.

Landeira, L. F. L., Paz, J. A. S., Cunha da Costa, R. M. S., Goes, L., Siqueira, J. D., Ferreira, S. C., Vizzoni, V. F., Caetano, R. da S., Siqueira, E., Cardoso, K. M., Ferreira, S. N., Silva, J. da, Miyashiro, L. M., Soares, M., Barroso, S. P. C., & Nico, D. (2025). [reference]. Detection of SARS-CoV-2 in rectal swabs from dogs and cats in São João de Meriti, Rio de Janeiro, Brazil. Ciência Rural, 55(11), e20240157.

Miragaya, M., Jolly, A., Carrizo, A., Cicuttin, G., Eiras, D. F., Bertuzzi, M., Jar, A. M., González López Ledesma, M. M., Gamarnik, A. V., Lucero Arteaga, F., Bucafusco, D., Capitelli, G., & Mundo, S. L. (2025). [reference]. SARS‑CoV‑2 in dogs: gamma variant and its transmission from humans in Ciudad de Buenos Aires, Argentina. InVet (Investigación Veterinaria), 27, 1‑8.

Ogoti, B. M., Riitho, V., Rodon, J., Mutono, N., Tesch, J., Oyugi, J., … Müller, M. A. (2025). [reference]. On‑site detection of MERS‑CoV infections in a camel slaughterhouse in Kenya using a commercial rapid antigen test. Frontiers in Veterinary Science, 12.

Ramasamy, S., Bustamante, F., LaBella, L. C., Cole, S. D., Armstrong, J., Thompson, D. L., Hardham, J. M., Kuchipudi, S. V., & Tewari, D. (2025). [reference]. Spillover of SARS-CoV-2 to domestic dogs in COVID-19-positive households: A one health surveillance study. Virus research, 360, 199629. Advance online publication.

Shi, L., Roy, S., Lang, Y., Wen, Y., Mitchell, W. J., Yang, W., Wang, L., Zhang, J., Liu, H., Driver, J. P., Peiris, M., & Ma, W. (2025). [reference]. A single‑dose intranasal immunization with a novel bat influenza A virus‑vectored MERS vaccine provides effective protection against lethal MERS‑CoV challenge. mBio, 16(8), e01107‑25.

Trifonova, I., Korsun, N., Alexiev, I., Ivanov, I., Grigorova, L., Stoikov, I., Donchev, D., Tchakarova, S., Lazarova, L., & Christova, I. (2025). [reference]. Tracking two-way human-mink transmission during an outbreak of severe acute respiratory syndrome coronavirus 2 on a farm in Bulgaria. Problems of Infectious Parasitic Diseases, 53(2).

Wang, H., Meng, Y., Chen, X., Cui, X., Zuo, Q., Han, N., Liang, X., Shen, X., Li, C., Li, D., Wang, F., He, L., Chen, R., Lu, X., You, W., Wu, A., Chen, R.‑A., Chen, W., Ding, C., & Shen, Y. (2025). [reference]. Unveiling novel viral diversity, biogeography, and host networks in wildlife through high‑throughput sequencing data mining. Advanced Science.

FAO actions

  • FAO, in collaboration with the World Health Organization (WHO) and the Coalition for Epidemic Preparedness Innovations (CEPI), will organize a two-day workshop on 18-19 November 2025 entitled Use-case workshop for Middle East Coronavirus Vaccines. The workshop will include a One Health session to explore the cost-benefits of using MERS vaccines in camels to protect public health.
  • FAO, WHO, and WOAH hosted two webinars through the Global MERS community on 18 and 25 March 2025. FAO experts presented the FAO recommendations for camel sampling, the investigation of MERS-CoV, SARS-CoV-2 and other coronavirus recombination in camels, along with the results of the work conducted in Ethiopia, Kenya and Oman.
  • FAO, WHO and the University of Calgary launched the MERS-Tracker: An interactive dashboard to support evidence-based decision making. It consists of a data tracking platform and a dashboard displaying published MERS-CoV serosurveys, viral testing, and genomic sequencing studies in animals and humans. The dashboard was launched during a WHO EPI-WIN webinar and is accessible here: MERS-Tracker.
  • In April 2024, FAO published a highlight paper for the International Year of Camelids 2024 entitled “Dromedary camels and MERS-CoV: filling knowledge gaps”.
  • The coronavirus network (COVINET) meeting was held from 26 to 27 March 2024 in Geneva, Switzerland. FAO and representatives from the reference centers on zoonotic coronaviruses attended the meeting.
  • FAO, WHO, WOAH and UNDP have organized the Quadripartite Global Technical Meeting on MERS-CoV and Other Emerging Zoonotic Coronaviruses, held from 27-29 November 2023 in Riyadh, Kingdom of Saudi Arabia.
  • FAO takes part in regular WHO virus evolution group meetings to discuss latest findings on SARS-CoV-2 variants of interest and variants of concern.
  • FAO has designated FAO Reference Centres for Zoonotic Coronaviruses. To date, Istituto Zooprofilattico Sperimentale delle Venezie (IZSVe) and Istituto Zooprofilattico Sperimentale dell'Abruzzo e del Molise Giuseppe Caporale (IZSAM), Italy (joint center), the Federal State-Financed Institution “The Federal Center for Animal Health” (FSVPS-FGBI ARRIAH), Russian Federation , CSIRO Australian Centre for Diseases Preparedness (ACDP), Australia, and Ohio State University Infectious Diseases Institute (OSU-IDI), USA, Institut de recherche pour le développement (IRD) and Centre de coopération internationale en recherche agronomique pour le développement (CIRAD) of France (joint center), and Ministry for Primary Industries, New Zealand have been appointed. Furthermore, Friedrich-Loeffler-Institut (FLI),Germany designated as Reference Centre for Emerging Zoonotic Pathogens and High Biosecurity/ Biocontainment Facilities.
  • The Joint FAO/IAEA Centre (Nuclear Techniques in Food and Agriculture) (CJN) is working, through its veterinary laboratory network in 69 countries, to support diagnosis of SARS-CoV-2 in animals and monitoring.

FAO publications

Next issue: December 2025
The disease situation updates are produced by the FAO Emergency Prevention System for Animal Health (EMPRES-AH) as part of its mission to increase global disease intelligence.
Disclaimer

Information provided herein is current as of the date of issue. Information added or changed since the last SARS-COV-2 animal situation update appears in orange. Human cases are depicted in the geographic location of their report. For some cases, exposure may have occurred in one geographic location but reported in another. For cases with unknown onset date, reporting date was used instead. FAO compiles information drawn from multiple national (Ministries of Agriculture or Livestock, Ministries of Health; Centers for Disease Prevention and Control [CDC]) and international sources (World Health Organization [WHO], World Organisation for Animal Health [WOAH]) as well as peer-reviewed scientific articles nd preprints. FAO makes every effort to ensure, but does not guarantee, accuracy, completeness or authenticity of the information. The boundaries and names shown and the designations used on these map(s) do not imply the expression of any opinion whatsoever on the part of FAO concerning the legal status of any country, territory, city or area or of its authorities, or concerning the delimitation of its frontiers and boundaries. Dashed lines on maps represent approximate border lines for which there may not yet be full agreement.

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